Кавказский энтомол. бюллетень 13(2): 161–233 © CAUCASIAN ENTOMOLOGICAL BULL. 2017


The longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) in the fauna of Asia.

  1. New or little-known taxa, mainly from Indochina and Borneo, with reviews of some genera

    Жуки-дровосеки трибы Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) фауны Азии.

    1. Новые и малоизвестные таксоны, преимущественно из Индокитая и Борнео, с обзорами некоторых родов

A.I. Miroshnikov1, 2

А.И. Мирошников1, 2

1Russian Entomological Society, Krasnodar, Russia. E-mail: miroshnikov-ai@yandex.ru

2Sochi National Park, Moskovskaya str., 21, Sochi, Krasnodar Region 354002 Russia

1Русское энтомологическое общество, Краснодар, Россия

2Сочинский национальный парк, ул. Московская, 21, Сочи, Краснодарский край 354002 Россия

Key words: Coleoptera, Cerambycidae, Cerambycini, new genera and species, reviews of genera, Southeastern Asia.

Ключевые слова: Coleoptera, Cerambycidae, Cerambycini, новые роды и виды, обзоры родов, Юго-Восточная Азия.


Abstract. Taking into account the new data presented in this paper, the tribe Cerambycini in the fauna of Asia contains almost 60 genus-group taxa and about

335 species, being the largest compared to the faunas of the other parts of the world. Serious problems in the development of a reasonable supraspecific classification of the tribe are noted. Reviews of such taxonomically confused genera as Elydnus Pascoe, 1869, Imbrius Pascoe, 1866, Zatrephus Pascoe, 1857, and Zegriades Pascoe, 1869, as well as keys to their constituent species are given. The following new genera and species are described, two new statuses change and new combinations are established: Falsopachydissus gen. n., F. foveiscapus (Holzschuh, 2011), comb. n., Mimimbrius gen. n., M. dembickyi sp. n. (Southern Thailand), M. geminatus (Holzschuh, 2005), comb. n., M. micaceus (Pascoe, 1858), comb. n.,

M. subargenteus (Gressitt et Rondon, 1970), comb. n.,

Pascoetrephus gen. n., P. hefferni sp. n. (Eastern Malaysia),

P. klimenkoi sp. n. (Eastern Malaysia), P. inscitus (Pascoe, 1857), comb. n., P. ranongensis (Holzschuh, 2009), stat. n. et comb. n., Spinidymasius gen. n., S. chrysophanes (Gressitt et Rondon, 1970), comb. n., S. crinicornis (Hüdepohl, 1989), comb. n., S. dembickyi (Holzschuh, 2003), comb. n.,

S. grossescapus (Hüdepohl, 1989), comb. n., S. huedepohli (Vives, 2005), comb. n., S. ochraceovittatus (Hüdepohl, 1989), comb. n., S. pascoei (Gahan, 1891), comb. n.,

S. sericatus (Pascoe, 1869), comb. n., S. tawauanus (Vives et Heffern, 2016), comb. n., Elydnus rufulus Holzschuh, 2016, stat. n. These new genera are also reviewed, with keys to their species given. The following new species are described: Dymasius makarovi sp. n. (Western Malaysia), D. murzini sp. n. (Sri Lanka),

image

DOI: 10.23885/1814-3326-2017-13-2-161-233

Elydnus barclayi sp. n. (Southern Thailand and Western Malaysia), E. tatianae sp. n. (Vietnam), E. vitalii sp. n. (Vietnam), Imbrius fedorenkoi sp. n. (Southern Vietnam),

I. klimenkoi sp. n. (Eastern Malaysia), I. solodovnikovi sp. n. (Eastern Malaysia), Massicus ivani sp. n. (Eastern Malaysia), M. valentinae sp. n. (Western Malaysia), Sebasmia indochinensis sp. n. (Thailand and Vietnam), Zatrephus golovatchi sp. n. (Southern Vietnam), Zegriades olemehli sp. n. (Eastern Malaysia). The genus Sebasmia Pascoe, 1859 is reported from Indochina for the first time. New records of a number of species from other genera are given as well, thus one way or another expanding their distribution areas, sometimes very significantly so. Provisional considerations on the taxonomy of the genus Pachydissus Newman, 1838 and on the systematic position of Plavichydissus Pic, 1946 are presented. Based on a comparison of the holotypes of Dymasius macilentus (Pascoe, 1859) and D. strigosus J. Thomson, 1864, their synonymy traditionally used in the literature is questioned as requiring indisputable evidence. The lectotype male of Dymasius minor Gahan, 1906 is designated. Abundant pictures of the species studied, including numerous type specimens, are provided.

Резюме. Триба Cerambycini Latreille, 1802 с учетом новых данных, представленных в настоящей работе, насчитывает в фауне Азии почти 60 таксонов родовой группы и около 335 видов – наибольшее число таксонов по сравнению с другими частями света. Обсуждаются серьезные проблемы в построении обоснованной надвидовой классификации трибы. Даны обзоры таких таксономически запутанных родов, как Elydnus Pascoe, 1869, Imbrius Pascoe, 1866, Zatrephus Pascoe,


1857, Zegriades Pascoe, 1869, и предложены таблицы для определения их видов. Описаны следующие новые роды и виды и установлены новые комбинации и два новых статуса: Falsopachydissus gen. n., F. foveiscapus (Holzschuh, 2011), comb. n., Mimimbrius gen. n.,

M. dembickyi sp. n. (Южный Таиланд), M. geminatus (Holzschuh, 2005), comb. n., M. micaceus (Pascoe, 1858), comb. n., M. subargenteus (Gressitt et Rondon, 1970), comb. n., Pascoetrephus gen. n., P. hefferni sp. n. (Восточная Малайзия), P. klimenkoi sp. n. (Восточная Малайзия), P. inscitus (Pascoe, 1857), comb. n.,

P. ranongensis (Holzschuh, 2009), stat. n. et comb. n., Spinidymasius gen. n., S. chrysophanes (Gressitt et Rondon, 1970), comb. n., S. crinicornis (Hüdepohl, 1989), comb. n., S. dembickyi (Holzschuh, 2003), comb. n.,

S. grossescapus (Hüdepohl, 1989), comb. n., S. huedepohli (Vives, 2005), comb. n., S. ochraceovittatus (Hüdepohl, 1989), comb. n., S. pascoei (Gahan, 1891), comb. n.,

S. sericatus (Pascoe, 1869), comb. n., S. tawauanus (Vives et Heffern, 2016), comb. n., Elydnus rufulus Holzschuh, 2016, stat. n. Даны обзоры этих новых родов и таблицы для определения их видов. Описаны следующие новые виды: Dymasius makarovi sp. n. (Западная Малайзия), D. murzini sp. n. (Шри Ланка), Elydnus barclayi sp. n. (Южный Таиланд и Западная Малайзия), E. tatianae sp. n. (Вьетнам), E. vitalii sp. n. (Вьетнам), Imbrius fedorenkoi sp. n. (Южный Вьетнам), I. klimenkoi sp. n. (Восточная Малайзия),

I. solodovnikovi sp. n. (Восточная Малайзия), Massicus ivani sp. n. (Восточная Малайзия), M. valentinae sp. n. (Западная Малайзия), Sebasmia indochinensis sp. n. (Таиланд и Вьетнам), Zatrephus golovatchi sp. n. (Южный Вьетнам), Zegriades olemehli sp. n. (Восточная Малайзия). Род Sebasmia Pascoe, 1859 впервые приведен для Индокитая. Отмечены также новые находки целого

ряда видов из других родов, расширяющие их ареалы. Обсуждается таксономия рода Pachydissus Newman, 1838 и систематическое положение Plavichydissus Pic, 1946. На основании сравнения голотипов Dymasius macilentus (Pascoe, 1859) и D. strigosus J. Thomson, 1864 обосновано, что традиционно используемая в литературе синонимия D. macilentus = D. strigosus требует неоспоримых доказательств. Обозначен лектотип Dymasius minor Gahan, 1906. Представлено большое количество иллюстраций исследуемых видов, в том числе многих типовых экземпляров.

Introduction

Taking into account the new data presented in this paper, the Asian fauna of the tribe Cerambycini Latreille, 1802 contains almost 60 genus-group taxa and about 335 species, being the largest compared to the faunas of the other parts of the world.

The tribe is generally characterized by a very considerable morphological diversity, on the one hand, and by very similar or identical morphological structures in these or those component groups, on the other. All this makes it extremely difficult any clear differential diagnoses

of various genus-group taxa to be worked out and poses serious problems in the development of a reasonable supraspecific classification of the tribe.

An extensive range of questions pertaining to the systematics and morphology of various representatives of the tribe were recently discussed by the author at the 15th Congress of the Russian Entomological Society [Miroshnikov, 2017]. Many of the ideas expressed there have been implemented in one way or another in the present paper.

A detailed study of a highly rich and diverse material amassed by the author, partly through the invaluable assistance of numerous colleagues and friends, has allowed to reconsider the species composition and provide reviews of such taxonomically confused genera as Elydnus Pascoe, 1869, Imbrius Pascoe, 1866, Zatrephus Pascoe, 1857 and Zegriades Pascoe, 1869; to describe and revise four new genera; to discuss some aspects of the systematics of the genera Dymasius J. Thomson, 1864, Massicus Pascoe, 1867, Pachydissus Newman, 1838 and their individual representatives; to describe sixteen new species belonging to nine genera, including two new ones; to clarify or expand, partly very significantly, the distribution areas of a number of species; and to present some other new data on little-known forms.

No complete systematic list of the genus-group taxa

of Cerambycini is available in the modern literature. Nor even the exact number of cerambycine genera and species is known for the fauna of Asia. In the present paper, this information is not arranged in alphabetic order, but is given in a way that seems most convenient for a discussion and corresponding taxonomic conclusions. The species lists in the generic accounts are given, as far as possible, in systematic order.

I have properly remounted many of the type specimens kept in various museums in order to take high- quality photographs of both their habitus and individual taxonomically important details. Therefore, in the present work, the pictures of these or those types may differ to some degree in appearance from those available in previous publications, including original ones.

Given the various new records of these or those species generally from Indochina as presented in this paper, the Malay Peninsula, from where some of such records are known, is taken here as its southern extremity.

The material treated in this work belongs to the following institutional and private collections:

BM – Bishop Museum (Honolulu, USA);

BMNH – Natural History Museum (London, United Kingdom);

IRSN – Institut Royal de Sciences naturelles de Belgique (Bruxelles);

LIPI – Lembaga Ilmu Pengetahuan Indonesia (Cibinong-Bogor, Indonesia);

NHMD – Natural History Museum of Denmark, University of Copenhagen (Copenhagen, Denmark);

MNHN – Muséum national d’Histoire naturelle (Paris, France);

PUM – Moscow Pedagogical State University (Moscow, Russia);


USNM – National Museum of Natural History, Smithsonian Institution (Washington D.C., USA);

ZIN – Zoological Institute of the Russian Academy of Sciences (St. Petersburg, Russia);

ZSM – Zoologische Staatssammlung München (München, Germany);

cAM – collection of Alexandr Miroshnikov (Krasnodar, Russia);

cAN – collection of Alexandr Napolov (Riga, Latvia); cCH – collection of Carolus Holzschuh (Villach,

Austria);

cDH – collection of Daniel Heffern (Houston, USA); cEV – collection of Eduard Vives (Barcelona, Spain); cFV – collection of Francesco Vitali (Luxembourg); cGC – collection of Gérard Chemin (Champigny-sur-

Marne, France);

cLD – collection of Luboš Dembický (Brno, Czech Republic);

cNO – collection of Nobuo Ohbayashi (Kamimiyada, Miura City, Japan);

cPH – collection of Pierre Haller (Essert, Le Mouret, Switzerland);

cPJ – collection of Philippe Jacquot (Montboucher- sur-Jabron, France);

cSM – collection of Sergey Murzin (Moscow, Russia); cTT – collection of Tomáš Tichý (Opava, Czech

Republic);

cWT – collection of William Tyson (Coarsegold, USA).

Results and discussion

Tribe Cerambycini Latreille, 1802 Genus Imbrius Pascoe, 1866

Imbrius Pascoe, 1866b: 528. Pascoe, 1869: 518; Lacordaire,

1868: 261 (syn. pro Dymasius); Gemminger, 1872: 2803 (syn. pro

Dymasius); Aurivillius, 1912: 59; Hüdepohl, 1990a: 65; Heffern,

2013: 10.

Type species: Imbrius lineatus Pascoe, 1866, by subsequent designation [Pascoe, 1869].

Diagnosis. This genus differs from all related and other Asian genera of the tribe by the peculiar structure of the pronotum, namely, the combination of certain features of its shape, sculpture and setation, all being more or less similar in species of the genus, as in Figs 31–54.

The pronotum of Imbrius is clearly or at least slightly/ barely, but always longitudinal, more or less broadly rounded or obtusely angulate in the middle part on the sides, at the apex insignificantly narrower than at the base; the sculptural formations in the apical one-third are sharply delimited posteriorly by symmetrical, usually arcuate ledges creating two (very rarely three) protrusions near the midline, these usually being well-expressed, right- or obtuse-angled, sometimes sharpened and directed backwards, in front with a median, often longitudinal, more or less clear tubercle; the apex of the pronotum at the midline also has two symmetrical, more or less clear, usually right- or obtuse-angled, small protrusions or a single, transverse, truncate one; the remaining disc surface

covered with coarse, very coarse or more smooth tubercles and folds of various shapes, but is devoid of folds that are strongly extended in one direction or another; with a more or less strongly developed setation forming separate and different shapes, but neither long, sometimes partly fused, symmetrical bundles nor spots, often missing almost entirely or mostly on the disc, especially in its middle area; the setation never forms any obvious bands all or almost all along the pronotum.

Imbrius is also characterized by the following features, the combination of which makes it even more distinct compared to the other genera of the tribe: head comparatively short, usually with a deep, broad, longitudinal groove both between dorsal lobes of eyes and on vertex; eyes well-developed, more or less strongly convex; antennae of male different both in length and structure of antennomeres, sometimes much longer than body and with many antennomeres strongly elongated, but in most cases moderately long, with less strongly elongated antennomeres; male antennomeres 3–5 or 3–4 in the vast majority of cases more or less inflated in apical parts, length ratios of antennomeres differing, antennomeres 1 and 3–5 can be comparatively short and insignificantly different in length, antennomeres 6–10 moderately broadened towards apex, their apical inner angle sometimes with a clear spine, but thereby last antennomere apically of usual structure, not extended into a sharp spine; pronotum generally rounded or obtusely angulate in outline on sides, but can show separate protrusions formed by a very coarse sculpture, yet without clear lateral tubercles; elytra moderately, sometimes strongly elongated, without distinct longitudinal ribs, with both a relatively rough and very small, double, very contrastingly differing puncturation, as in Figs 64–66, apical sutural angle usually more or less straight, but sometimes sharpened or produced into a long tooth; recumbent setation of elytra entirely or mostly monochrome, not velvety, continuous (may be partly abraded) or, behind middle, with a v-shaped fascia devoid of dense setae (sometimes appearing like abraded fragments of setation), also can be in the form of more or less clear longitudinal strips; prosternum with a well- expressed transverse groove in front of middle, prosternal process without lateral dentiform protrusion on sides in middle part of apical half, as in Figs 73–78, usually with a more or less clear, sometimes very sharp tubercle, but can also be without it; mesosternal process without tubercle, between coxae clearly or slightly wider than prosternal process, sometimes these processes subequal in width between coxae; legs moderately long, femora claviform, both femora and tibiae without carina along each side; genitalia as in Figs 82–110, endophallus of various shapes, thereby can differ very strongly between superficially very similar species (Figs 102–105; see also below); body predominantly medium-sized, 13.5–26.5 mm in length.

Besides this, some important details of Imbrius

structure are shown below, in the diagnosis of the new genus, Mimimbrius gen. n., described here.

Composition. The genus includes twelve species, three of which are described as new.

Disribution. Oriental realm.


image

Figs 1–6. Imbrius Pascoe, 1866, habitus, dorsal view.

1–2 – I. lineatus Pascoe, 1866; 3, 6 – I. similis Hüdepohl, 1990; 4–5 – I. solodovnikovi sp. n. 1–2 – syntypes; 3–4 – holotypes; 5 – paratype; 1, 3–4 –

males; 2, 5–6 – females.

Рис. 1–6. Imbrius Pascoe, 1866, общий вид сверху.

1–2 – I. lineatus Pascoe, 1866; 3, 6 – I. similis Hüdepohl, 1990; 4–5 – I. solodovnikovi sp. n. 1–2 – синтипы; 3–4 – голотипы; 5 – паратип; 1, 3–4 –

самцы; 2, 5–6 – самки.


Imbrius lineatus Pascoe, 1866 (Figs 1, 2, 35–38, 41, 42, 64, 69, 73–75,

84, 85, 89, 90, 435, 436)

Imbrius lineatus Pascoe, 1866b: 529, pl. 41, fig. 12. Type locality: [Malaysia] Penang (according to the original description and the label of the holotype). Aurivillius, 1912: 60 (Penang); Hüdepohl, 1990a: 70 (Malaysia; Sumatra).

Dymasius lineatus: Gemminger, 1872: 2803 (Penang).

Material. ♂, syntype, (BMNH) (Fig. 1), “Penang (Lamb.) Pascoe Coll.”, “Pascoe Coll. 93–60”, “Imbrius lineatus Pasc. ♂ Type”, “Type”, “♂”, “Imbrius lineatus P. Type Penang” (Fig. 435); ♀, syntype, (BMNH) (Fig. 2), “Penang”, “Penang (Lamb.) Pascoe Coll.”, “Pascoe Coll. 93–60”, “Imbrius lineatus Pasc.

♀ Type”, “Type”, “♀” (Fig. 436); 1♂ (BMNH), “Singapore”, “Atkinson Coll. 92–3”, “Imbrius lineatus Pascoe, 1866 ♂ det. A. Miroshnikov 2017”; 1♀ (BMNH), “Malay, Penang”, “Fry Coll. 1905.100”, “16724”, “Imbrius lineatus Pasc., ♀ comp. with type”; 1♀ (ZSM), “N Sumatra, Aik Tarum, 1.I.[19]84 Diehl [leg.]”, “Imbrius lineatus Pasc. Hüdepohl det. 1987”.

Morphological notes. Body length 13.5–15.8 mm, humeral width 3.1–3.8 mm, thereby syntype male largest.

Disribution. Western Malaysia, Singapore, Indonesia (Sumatra).

Imbrius solodovnikovi Miroshnikov, sp. n.

(Figs 4, 5, 31–33, 39, 40, 65, 68, 77, 82, 83, 87, 88)

Imbrius lineatus auct.: Heffern, 2013: 10 (partim, Borneo) (non Pascoe, 1866).

Material. Eastern Malaysia: holotype, ♂ (NHMD) (Fig. 4): Sabah, Trus

Madi Mt., 03.2004 (local collector), “Imbrius lineatus Pascoe, Ole Mehl det. 2005”. Paratypes: 3♀ (NHMD), same locality as holotype, 03.2003, 03.2004, 04.2005 (local collector), “Imbrius lineatus Pasc., Ole Mehl det. 2005”; 1♀ (cPJ), same locality as holotype, 1200 m, 13–26.05.2017 (leg. A. Sochivko); 1♀ (BMNH), “Sarawak, R. Kapah trib., of R. Tinjar, 3.X.1932”, “Beating”, “Primitive forest”, “Oxford Univ. Exp. B.M. Hobby & A.W. Moore, B.M.1933– 254”; 1♀ (BMNH), same labels, “Elydnus lineatus Pasc.”; 1♀ (cDH), Sabah, Trus Madi Mt., 1500–2000 m, 11.03.1998 (leg. Affenddy), “Imbrius lineatus Pascoe, det. K. Hüdepohl [19]99”; 1♀ (cDH), same geographical label, “Imbrius lineatus Pascoe, det. D. Heffern”; 1♀ (cDH), Sabah, Trus Madi Mt., 1500–2000 m, 03–05.1998 (local collector), “Imbrius lineatus Pascoe, det. K. Hüdepohl [19]99”; 1♀ (cDH), same geographical label, “Imbrius lineatus Pascoe, det. D. Heffern”; 1♀ (cLD) (Fig. 5), Sabah, Batu Punggul Resort env., primary forest, 07.1999 (leg. J. Kodada); 1♀ (cAM ex NHMD), 1♀ (NHMD), Sabah, Crocker Range, 03.2004, 04.2005 (local collector), “Imbrius lineatus Pasc., Ole Mehl det. 2005”; 1♂ (BMNH), “Borneo, Sabah, Ranau, 24.IX.2005, Steven Chew [leg.], BMNH {E}2006-36”, “Imbrius lineatus Pasc., det. C. Holzschuh 2009”. 2♀ (NHMD), 1♀ (cAM ex NHMD), Sabah, Crocker Range, 04.2014 (local collector), “Imbrius lineatus Pasc., O. Mehl det. 2014”; 2♀ (cAM), Sabah, Trus Madi Mt., 1250 m, 05°26’35”N / 116°27’5”E, 22–26.05.2014, 5–12.11.2015 (leg. A. Klimenko).

Diagnosis. This new species is very similar to I. lineatus, but differs by the structure of the male genitalia, in particular, the penis being relatively wide near the apex, narrowed at the very apex through a short cone, as in Figs 82, 83 (cf. Figs 84, 85); by the noticeably wider parameres, in general the peculiar shape of the apical part of the tegmen, the wider base of the tegmen, as in Figs 87, 88 (cf. Figs 89, 90); by the more strongly developed, recumbent, light setation of the pronotum disc, as in Figs 31–33 (cf. Figs 35–38); the usually less strongly expressed, median, longitudinal tubercle near the apex of the pronotum, as in Figs 31–33 (cf. Figs 35–38); the somewhat denser, recumbent, light setation of the prosternum between the constriction at its apex and the anterior border of the coxal cavities, as in Figs 39, 40 (cf. Figs 41, 42); the usually denser setation of the apical part of the mesosternum, and often by the less clearly expressed longitudinal strips of dense, recumbent, light setae on the elytra. Imbrius solodovnikovi sp. n. can

also be compared to I. similis Hüdepohl, 1990, but differs clearly by the structure of the pronotum, in particular, its shape being slightly more longitudinal, the coarser sculpture, at least so on its disc, and the usually more or less clearly bidentate median protrusion at the apex; if longitudinal strips of dense, recumbent, light setae on the elytra clearly expressed, then these strips being narrower; the sharper apical tubercle of the prosternal process; the weakly or barely impressed profemora both on the external and inner sides near the apex; the structure of the male genitalia (cf. Figs 86, 91).

Description. Body length 14.9–22.5 mm, humeral width 3.6–

5.2 mm, thereby holotype 17.4 and 4 mm, respectively. Coloration of integument mainly combines reddish brown and dark reddish brown tones; eyes, partly mandibles, head dorsally or, besides this, partly pronotum black.

Head with well-developed antennal tubercles; with a sharp median groove between bases of antennae, growing even sharper between eyes and on vertex, as well as with a very sharp, deep, oblique groove behind inner part of each of dorsal lobes of eyes; genae rather short; eyes well-developed, relatively strongly convex; submentum in basal part with a coarse irregular sculpture, in apical part strongly impressed, smooth; neck with sharp transverse folds both ventrally and laterally; antennae of both male and female usually subequal in length, reaching beyond apex of elytra by last antennomere, but in female sometimes can only reaching the apex of elytra; length ratio of antennomeres 1–11 in male (holotype taken as an example), 30 : 10 : 30 : 26 : 28 : 37 : 41 : 42 : 40 : 37 : 48, in female (one of the paratypes taken as an example), 31 : 10 : 31 : 24 : 27 : 38 : 42 : 41 : 40 : 38 : 52; antennomere 1 with a heterogeneous, partly rough, rugose, dense puncturation; antennomere 2 subequal in length and width; male antennomeres 3 and 4 strongly, 5th moderately inflated in apical part; last antennomere with a distinct appendage.

Pronotum clearly longitudinal, 1.17–1.21 times as long as width; base 1.10–1.15 times as wide as apex; with a sharp constriction near apex; on disc with coarse or rough tubercles and folds, at apex in middle part with two more or less right-angled, small protrusions, as in Figs 31–33; near apex with a clearly, but not too strongly expressed, usually moderately shiny, longitudinal, median tubercle usually bearing at least a few, recumbent, light setae, as in Figs 31–33 (in I. lineatus, median tubercle strongly expressed, very shiny, completely devoid of recumbent light setae, as in Figs 35–38).

Scutellum strongly narrowed towards apex, triangular, at the very apex more or less broadly rounded or truncate, but can also be sharpened.

Elytra in male slightly narrowed towards apex, in female about parallel-sided, 2.67–2.88 times as long as humeral width; with both a relatively rough and very small, double, very contrastingly differing puncturation characteristic of the genus; apical external angle broadly or obtusely rounded, sutural angle narrowly rounded or obtuse.

Prosternum with a well-expressed, pretty deep transverse grooveinfrontofmiddle, withheterogeneous, coarseorroughfolds; prosternal process with a very clear apical tubercle; mesosternal process without tubercle dorsally, between coxae slightly or barely wider than prosternal process; mesosternum partly, metasternum and sternites with a small dense puncturation; metasternum with a clear, but not too sharp, median groove; last (visible) sternite at apex in male with a very clear broad emargination, in female widely rounded; last (visible) tergite at apex in male truncate, in female widely rounded.

Legs moderately short; femora claviform; tarsomere 1 very clearly shorter than tarsomeres 2 and 3 combined.

Recumbent setation mainly yellow tones, only partly greyish or greyish yellow; distribution of setation predominantly almost


same as in I. lineatus, but on disc of pronotum comparatively more strongly developed, thereby on tubercles lateral to midline, especially in basal part, always with clear fragments of setae while a fascia near apex less clearly interrupted in area of median tubercle, as in Figs 31–33 (in I. lineatus, tubercles lateral to midline devoid of setae, as a rule, only rarely with a few setae on tubercles in basal part just beyond middle; fascia near apex usually very clearly interrupted in area of median tubercle, as in Figs 35–38), on elytra often forming less strongly expressed, partially confused, longitudinal strips, as in Figs 4, 5, on prosternum partly denser, as in Figs 39, 40; head, pronotum laterally in apical part, pro- and metasterna partly, abdomen usually at apex, most of antennomeres in apical part, legs mainly on trochanters with more or less long, erect, partly suberect, sparse or individual, thin setae.

Genitalia of male as in Figs 82, 83, 87, 88.

Etymology. I am pleased to dedicate this new species to my colleague and friend, Dr. Alexey Yu. Solodovnikov, curator of the Coleoptera collection of the Natural History Museum of Denmark (University of Copenhagen), who constantly provides his great and versatile help to my research. Without his strong support, many of the issues, including those related to a rapid and safe delivery of material, would have been much more difficult to solve.

Disribution. Eastern Malaysia.

Imbrius similis Hüdepohl, 1990 (Figs 3, 6, 34, 86, 91, 437)

Imbrius similis Hüdepohl, 1990a: 70. Type locality: Philippines (according to the original description and the label of the holotype).

Material. ♂, holotype, (ZSM) (Fig. 3), “Philippines”, “Holotypus

♂ Imbrius similis mihi, Hüdepohl 1987” (Fig. 437); 1♀ (BMNH) (Fig. 6), “Phillip [sic; = Philippines] Islands”, “Butuan”, “Fry Coll. 1905.100”, “Imbrius similis Hüdepohl, 1990 ♀ det. A. Miroshnikov 2017”.

Morphological notes. This species was described from a single male which I have revised, its body length being 15.7 mm and humeral width 4 mm. I have also examined one female from BMNH with a body length of

14.5 mm and a humeral width of 3.65 mm.

Disribution. Philippines.

Imbrius ephebus Pascoe, 1866 (Figs 15, 18, 47, 441)

Imbrius ephebus Pascoe, 1866b: 529. Type locality: Java (according to the label of the holotype) (see Remarks). Pascoe, 1869: 519 (Singapore, Java, Penang); Aurivillius, 1912: 59 (Malacca, Java); Heffern, 2013: 10 (Borneo, Java, Western Malaysia).

Dymasius ephebus: Gemminger, 1872: 2803 (Penang).

Imbrius imitator Holzschuh, 2006: 222 (Malaysia, Sabah, Crocker Range). Holzschuh, 2010: 149 (syn. pro Imbrius ephebus). Material. ♀, holotype, by monotypy (see Remarks) (BMNH) (Fig.

15), “Java”, “Imbrius ephebus Pasc. Type”, “Type”, “Imbrius ephebus P. Java” (Fig. 441); 1♀ (BMNH), “Sarawak” (upperside), “♀” (underside); 1♀ (BMNH), “Sarawak”, “♀”; 1♀ (BMNH), “Sarawak, 76.24”, “Imbrius ephebus Pasc. ♀”; 1♀ (BMNH), “Borneo”, Fry Coll. 1905.100.”, “46359”, “Imbrius ephebus Pasc.

♀, comp. with type”; 1♀ (BMNH), “[Java] Noesa Cambangan, 23.IV.1932,

F.C. Drescher [leg.]”, “Brit. Mus. 1937–662”, “B.M. F.C.D. 1”; 1♀ (IRSN), “Java”; 1♀ (BMNH), “Sing[apore]”, “Pascoe Coll. 93–60”, “Imbrius ephebus Pascoe, 1866 ♀ det. A. Miroshnikov 2017”; 1♀ (NHMD), E Malaysia, Sabah, Crocker Range, 04.2014 (local collector), “Imbrius simulator Holz., Ole Mehl det. 2014”, “Imbrius ephebus Pascoe, 1866 ♀ det. A. Miroshnikov 2017”; ♂, holotype of Imbrius imitator Holzschuh, 2006 (cCH) (photograph); Fig. 18).

Morphological notes. Body length 16.5–19.8 mm, humeral width 4.1–4.8 mm, thereby holotype 19.3 and

4.7 mm, respectively.

Remarks. This species was described in the following work: “Catalogue of Longicorn Coleoptera collected in the Island of Penang by James Lamb, Esq. (Part II.)” [Pascoe, 1866b]. However, BMNH keeps the female type bearing 4 labels: “Java”, “Imbrius ephebus Pasc. Type”, “Type” and “Imbrius ephebus P. Java” (Fig. 441). Its length is 19.3 mm (humeral width 4.7 mm), which corresponds approximatelly to the length indicated in the original description (9 lines,

i.e. about 19.05 mm). I am not aware of any other types.

In a later work, Pascoe [1869: 519] noted the following distribution area of this species: “Singapore, Java (and Penang)”. However, the material from “Penang”, as well as those from other districts of Western Malaysia, are known from neither BMNH nor any other of the museums whence material was possible to study.

Therefore, taking into account all above, it seems obvious that Pascoe provided the original description, based exactly on the specimen kept in BMNH, but for some reason he falsely indicated its provenance from “Penang”; the same holds true also for the introductory text of the first part of the work [Pascoe, 1866a]. In this respect, I consider that specimen as the holotype by monotypy, and I define the type locality according to the appropriate label (“Java”), but not to the original description (“Penang”).

Disribution. Western and Eastern Malaysia, Indonesia (Java), Singapore.


Imbrius diehli Hüdepohl, 1989 (Figs 13, 14, 16, 17, 48, 49, 67, 442, 443)

Imbrius diehli Hüdepohl, 1989a: 48. Type locality: [Indonesia] Sumatra, Aik Tarum (according to the original description and the label of the holotype). Heffern, 2013: 10 (Borneo, Sumatra).

Material. ♂, holotype, (ZSM) (Fig. 16), “N–Sumatra, Aik Tarum,

1.I.[19]84, Diehl [leg.]”, “Holotypus ♂ Imbrius diehli mihi, Hüdepohl 1988” (Fig. 442); 1♀, paratype, (ZSM) (Fig. 17), “N–Sumatra, Dolok Merangir, VIII.[19]81, Diehl leg.”, “Paratypus ♀ Imbrius diehli mihi, Hüdepohl 1988” (Fig. 443); 1♂ (BMNH), “N. Borneo”, “Pascoe Coll. 93–60”, “♂”, “Imbrius diehli Hüdepohl, 1989 ♀ det. A. Miroshnikov 2017”; 1♂ (NHMD) (Fig. 13), E Malaysia, Sabah, Trus Madi Mt., 04.2014 (local collector), “Imbrius diehli Hüdepohl, 1989 ♂ det. A. Miroshnikov 2017”; 1♀ (BMNH) (Fig. 14), “Brunei: Temburong District, ridge NE of Kuala Belalong, 125W m. v. light”, “approx. 300 m alt. XI.1992, J.H. Martin coll., BM 1992–172”, “1751”, “Imbrius ephebus Pascoe, E. Vives det. 2006”, “Imbrius diehli Hüdepohl, 1989

♀ det. A. Miroshnikov 2017”.

Morphological notes. Body length 20.1–26.5 mm, humeral width 4.65–6 mm, thereby holotype 21.5 and

4.9 mm, paratype 22.9 and 5.4 mm, respectively.

Disribution. Eastern Malaysia, Brunei, Indonesia (Sumatra).


Imbrius simulator Holzschuh, 2005 (Figs 20, 21, 44–46, 93, 94, 96, 97)

Imbrius simulator Holzschuh, 2005: 7. Type locality: Malaysia, Sabah, Crocker Range (according to the original description). Heffern, 2013: 10.

Material. ♂, holotype (cCH) (photograph); 1♂, paratype (cDH)

(Fig. 20), E Malaysia, Sabah, Crocker Range, 13.04.1999 (local collector), “Paratypus Imbrius simulator n. sp. det. C. Holzschuh 2004”; 1♂, 2♀ (NHMD), same locality, 03.2003, 03.2004 (local collector), “Imbrius simulator Holz., Ole Mehl det. 2005”; 1♂ (cAM ex NHMD), 2♀ (NHMD), 1♀ (cAM ex NHMD), E Malaysia, Sabah, Trus Madi Mt., 03.2003 (local collector), “Imbrius simulator Holz., Ole Mehl det. 2005”; 1♂, 2♀ (cAM) (Fig. 21), Sabah, Trus Madi Mt., 5.07.2011, 24.08.2012, 5.04.2013


image

Figs 7–12. Imbrius Pascoe, 1866, habitus, dorsal view.

7–8 – I. uniformis Holzschuh, 2010 (7 – after Holzschuh [2010], photograph by Luboš Dembický); 9, 12 – I. corrugatus Hüdepohl, 1990; 10–11 –

I. klimenkoi sp. n. 7, 9–10 – holotypes, males; 8 – female from Western Malaysia (Perak, Ringlet); 11–12 – paratypes, females. Рис. 7–12. Imbrius Pascoe, 1866, общий вид сверху.

7–8 – I. uniformis Holzschuh, 2010 (7 – по [Holzschuh, 2010], фотография Л. Дембицкого); 9, 12 – I. corrugatus Hüdepohl, 1990; 10–11 –

I. klimenkoi sp. n. 7, 9–10 – голотипы, самцы; 8 – самка из Западной Малайзии (Перак, Ринглет); 11–12 – паратипы, самки.


image

Figs 13–18. Imbrius Pascoe, 1866, habitus, dorsal view.

13–14, 16–17 – I. diehli Hüdepohl, 1989; 15, 18 – I. ephebus Pascoe, 1866 (18 – holotype of I. imitator Holzschuh, 2006, after Holzschuh [2006], photograph by Luboš Dembický). 13 – male from Eastern Malaysia; 14 – female from Brunei; 15–16 – holotypes, female and male, respectively; 17 – paratype, female.

Рис. 13–18. Imbrius Pascoe, 1866, общий вид сверху.

13–14, 16–17 – I. diehli Hüdepohl, 1989; 15, 18 – I. ephebus Pascoe, 1866 (18 – голотип I. imitator Holzschuh, 2006, по [Holzschuh, 2006], фотография Л. Дембицкого). 13 – самец из Восточной Малайзии; 14 – самка из Брунея; 15–16 – голотипы, самка и самец, соответственно; 17 – паратип, самка.


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Figs 19–24. Imbrius Pascoe, 1866, habitus, dorsal view.

19 – I. fedorenkoi sp. n.; 20–21 – I. simulator Holzschuh, 2005; 22–23 – I. acutipennis (Fisher, 1935); 24 – I. allardi Hüdepohl, 1992. 19, 24 – holotypes,

males; 20 – paratype, male; 21, 23 – females; 22 – male.

Рис. 19–24. Imbrius Pascoe, 1866, общий вид сверху.

19 – I. fedorenkoi sp. n.; 20–21 – I. simulator Holzschuh, 2005; 22–23 – I. acutipennis (Fisher, 1935); 24 – I. allardi Hüdepohl, 1992. 19, 24 – голотипы,

самцы; 20 – паратип, самец; 21, 23 – самки; 22 – самец.


image

Figs 25–30. Mimimbrius gen. n., habitus.

25–26 – M. micaceus (Pascoe, 1858), comb. n.; 27 – M. geminatus (Holzschuh, 2005), comb. n.; 28–29 – M. subargenteus (Gressitt et Rondon, 1970), comb. n.; 30 – M. dembickyi sp. n. 29–30 – holotypes, males; 25–27 – females; 28 – male from Northern Thailand; 25, 27–30 – dorsal view; 26 – ventral view.

Рис. 25–30. Mimimbrius gen. n., общий вид.

25–26 – M. micaceus (Pascoe, 1858), comb. n.; 27 – M. geminatus (Holzschuh, 2005), comb. n.; 28–29 – M. subargenteus (Gressitt et Rondon, 1970), comb. n.; 30 – M. dembickyi sp. n. 29–30 – голотипы, самцы; 25–27 – самки; 28 – самец из Северного Таиланда; 25, 27–30 – вид сверху; 26 – вид снизу.


(leg. A. Klimenko); 2♂ (NHMD), same locality, 04.2013 (local collector), “Imbrius simulator Holz., O. Mehl det. 2014”.

Morphological notes. Body length 13.9–19.2 mm, humeral width 3.3–4.5 mm.

Disribution. Eastern Malaysia.

Imbrius fedorenkoi Miroshnikov, sp. n.

(Figs 19, 43, 66, 76, 92, 95)

Material. Holotype, ♂ (ZIN) (Fig. 19): Vietnam, Lam Dong Prov., 25 km NNW of Bao Loc, Loc Bao env., 800 m, 11°44’18”N / 107°42’08”E, 5–20.04.2013 (leg. D. Fedorenko).

Diagnosis. This new species seems to be especially similar to I. simulator, but differs clearly by the structure of the tegmen, the shape of the apical part of the penis, as in Figs 92, 95 (cf. Figs 93, 94, 96, 97), as well as by the structure of the pronotum, in particular, the very poorly developed setation along its sides, the somewhat peculiar sculpture of the disc, as in Fig. 43 (cf. Figs 44–46), including the shape of the folds in the basal part near the midline, the generally broader and more rounded (not crest-like) folds dorsally, the predominantly clearly sharper puncturation of the folds, the generally more delicate, recumbent, light pubescence of the antennae, the coloration of the elytra with a distinctly creamy tone, and some other minor features.

Description. Male. Body length 17.1 mm, humeral width

4.3 mm. Coloration of integument mainly combines red-brown, dark red-brown and red tones; eyes, mandibles partly and pronotum on disc near apex lateral to median tubercle black.

Head with well-developed antennal tubercles; with a sharp median groove between bases of antennae, growing even sharper between eyes and on vertex, as well as with a very sharp, deep, wide, oblique groove behind inner part of each of dorsal lobes of eyes; genae moderately short; eyes well-developed, relatively strongly convex; submentum in basal part with a rough shallow puncturation, in apical part clearly impressed, smooth; neck with clear transverse folds both ventrally and laterally; antennae reaching beyond apex of elytra by last antennomere; length ratio of antennomeres 1–11, 28 : 9 : 31 : 27 : 28 : 31 : 38 : 41 : 40 : 37 : 45;

antennomere 1 with a heterogeneous, partly rough, rugose, dense puncturation; antennomere 2 subequal in length and width; antennomeres 3 and 4 clearly, 5th and 6th moderately inflated in apical part; last antennomere with a very evident appendage.

Pronotum distinctly longitudinal, 1.17 times as long as width, at base slightly wider than at apex; with a sharp constriction near apex; on sides angularly broadened towards the middle from both base and apex; on disc predominantly with coarse, partly rough tubercles and folds, at apex in middle part with a distinctly emarginate protrusion; near apex with a sharp, longitudinal, long, median tubercle; compared to I. simulator (as noted in Diagnosis partly), tubercles and folds mostly roundish dorsally, wider, with sharp punctures partly, thereby in basal part lateral to midline with two symmetrical, short, slightly sinuous, partly more or less roundish tubercles connected by a transverse, short, moderately rough fold and behind these tubercles with a median fold branched in basal part into two partly transverse short folds (while in I. simulator in apical part lateral to midline with two pairs of symmetrical relatively narrow folds, these often partially connected with each other, thereby upper pair clearly or very clearly coarser, strongly crest-shaped, whereas instead of lower pair of folds sometimes with one or even two transverse folds; besides this, median tubercle near apex usually shorter than in new species, sometimes even very short, somewhat roundish).

Scutellum strongly narrowed towards apex, triangular.

Elytra barely narrowed towards apex, 2.53 times as long as humeral width; with both a relatively rough and very small,

double, very contrastingly differing puncturation characteristic of the genus; apical external angle obtusely rounded, sutural angle obtuse.

Prosternum with a well-expressed transverse groove in front of middle, with a coarse sculpture between this groove and anterior border of coxal cavities; prosternal process with a very clear apical tubercle; mesosternal process without tubercle dorsally, between coxae distinctly wider than prosternal process; mesosternum partly, metasternum and sternites with a small dense puncturation; metasternum with a clear, but not too sharp, median groove; last (visible) sternite very broadly truncate at apex; last (visible) tergite with a barely visible emargination apically.

Legs relatively well-developed; femora claviform; tarsomere 1 very clearly shorter than tarsomeres 2 and 3 combined.

Recumbent setation on head dorsally and pronotum yellow tones, both on scutellum and elytra grey with cream tint (in I. simulator, recumbent setation of both scutellum and elytra grey tones, but without cream tint), on remaining parts grey tones; distribution of setation predominantly about same as in

I. simulator, on pronotum generally weakly developed, as in Fig. 43, fragments of setation on its disc present only in apical one- quarter and near base, but on sides almost absent (in I. simulator, pronotum on sides with well-expressed individual fragments of recumbent setation, as in Figs 44–46); on elytra continuous recumbent setation in apical part interrupted by a v-shaped broad fascia (in holotype, setation partly in area of fascia strongly abraded), as in Fig. 19; head, pronotum mainly on sides, near both apex and base on disc, the very base of elytra, prosternum, abdomen at apex, most of antennomeres in apical part, legs mainly on trochanters with more or less long, erect, partly suberect, sparse or individual, thin setae.

Genitalia as in Figs 92, 95.

Etymology. I am pleased to dedicate this new species to my colleague and friend, Dr. Dmitry N. Fedorenko (Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia), who has, well over a decade, collected in Vietnam a rich and very valuable material on Coleoptera, including cerambycids.

Disribution. Vietnam.

Imbrius uniformis Holzschuh, 2010 (Figs 7, 8, 50, 99, 101, 104, 105, 108–110)

Imbrius uniformis Holzschuh, 2010: 149. Type locality: Malaysia, Sabah, Crocker Range (according to the original description). Heffern, 2013: 10.

Material. ♂, holotype (cCH) (photograph; Fig. 7); 1♀, paratype

(see Remarks) (BMNH), “[W Malaysia] Pahang, F.M.S., March 1923, M.R. Henderson [leg.]”, “1937.425.”, “317”, “Paratypus Imbrius uniformis n. sp. det. Holzschuh 2010”; 1♀ (cAM ex cLD) (Fig. 8), W Malaysia, Perak, 40 km SE Ipoh, Banjaran Titi Wangsa, Ringlet, 900 m, 26–31.03.2000 (leg. P. Čechovský), “Imbrius uniformis Holzschuh, 2010 ♂ det. A. Miroshnikov 2017”; 1♂ (cLD), same labels, but taken on 25.03–3.04.2002; 1♂ (cAM ex cLD), W Malaysia, Johor, Endau – Rompin, Selendang, 1–4.03.1997 (leg. I. Jeniš), “Imbrius uniformis Holzschuh, 2010 ♂ det. A. Miroshnikov 2017”; 1♀ (cLD), W Malaysia, Pahang, 30 km SE Ipoh, Cameron Highlands, Tanah Rata, 1500 m, 14–17.03.1998 (leg. P. Čechovský), “Imbrius uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”; 1♀ (cAM), same locality, 04.2015 (local collector), “Imbrius uniformis Holzschuh, 2010 ♂ det. A. Miroshnikov 2017”; 1♀ (NHMD), E Malaysia, Sabah, Trus Madi Mt., 8.03.2000 (local collector), “Imbrius ephebus Pascoe, 1866, Ole Mehl det. 2011”, “Imbrius uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”; 2♀ (NHMD), same locality, 04.2010, 04.2014 (local collectors), “Imbrius uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”; 1♀ (cAM), same locality, 24.08.2012, 1160 m (leg. A. Klimenko), “Imbrius uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”; 2♀ (NHMD), E Malaysia, Sabah, Crocker Range, 03.2005, 07.2007 (local collector), “Imbrius ephebus Pascoe, 1866, Ole Mehl det. 2011”, “Imbrius uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”.


Comparative material. 2♀ (NHMD), E Malaysia, Sabah, Crocker Range, 03.2001, 03.2003 (local collectors), “Imbrius ephebus Pascoe, 1866, Ole Mehl det. 2011”, “Imbrius ?uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”; 1♀ (cAM), E Malaysia, Sabah, Trus Madi Mt., 1250 m, 05°26’35”N / 116°27’5”E, 5–12.11.2015 (leg. A. Klimenko), “Imbrius

?uniformis Holzschuh, 2010 ♀ det. A. Miroshnikov 2017”.

Morphological notes. Body length 15.6–23.3 mm, humeral width 3.6–5.7 mm.

Remarks. The original description [Holzschuh, 2010: 149] erroneously stated the paratype in the BMNH to be a male (“1 ♂ Pahang... (BMNH)”), but in fact it is a female.

Disribution. Western and Eastern Malaysia.

Imbrius klimenkoi Miroshnikov, sp. n.

(Figs 10, 11, 98, 100, 102, 103, 106, 107)

Material. Holotype, ♂ (cAM) (Fig. 10): E Malaysia, Sabah, Trus Madi Mt., 1160 m, 20.03.2012 (leg. A. Klimenko). Paratypes: 2♂, 1♀ (Fig. 11) (cAM), same label as holotype; 1♀ (cAM), same label as holotype, but taken on 27.02.2014; 1♂ (NHMD), Trus Madi Mt., 03.2004 (local collector), “Imbrius ephebus Pascoe, 1866, Ole Mehl det. 2011”; 1♀ (NHMD), E Malaysia, Sabah, Crocker Range, 03.2004 (local collector), “Imbrius ephebus Pascoe, 1866, Ole Mehl det. 2011”; 1♂ (cAM), Sabah, Trus Madi Mt., 1250 m, 05°26’35”N / 116°27’5”E, 22–26.05.2014 (leg. A. Klimenko); 2♂ (cAM), same label, but taken on 5–12.11.2015; 2♂ (cAM), 1♂ (BMNH ex cAM), E Malaysia, Sabah, Nabawan Distr., ~7 km N Pensiangan, 530 m, 04°35’16”N / 116°19’27”E, 27–31.05.2014 (leg. A. Klimenko).

Diagnosis. Based on external features of the male, this new species is very similar to I. uniformis, while their females seem to be identical. However, I. klimenkoi sp. n. differs at least by the structure of the antennae of the male, in particular, the less strongly inflated antennomeres 3–5 in the apical part, the more numerous, long, erect or suberect setae on the inner side of antennomeres 3–7 or 3–8 and, in most cases, by the somewhat shorter antennae. The most important differences between I. klimenkoi sp. n. and I. uniformis lie both in the shape and fine armature of the endophallus, as in Figs 102, 103 (cf. Figs 104, 105), as well as in the conformation of the penis and tegmen. In the new species, the apical part of the penis is wider, as in Fig. 98 (cf. Fig. 99), that of the tegmen is also wider and less strongly elongated, as in Fig. 100 (cf. Fig. 101), each of the parameres on the ventral side distal to about 3/4 is more or less strongly sloping down, thereby the brown coloration of the parameres noticeably or very clearly fails to reach their bases, as in Fig. 100, whereas in I. uniformis the parameres are more or less strongly sloping down only behind their bases on their ventral side so that the entire surface of the paramers lies in about the same plane, thereby the brown coloration covers completely each of the parameres and reaches their bases, as in Fig. 101. Reliable differences between I. klimenkoi sp. n. and I. uniformis have been found yet neither in the structure of the female genitalia nor in general any external features (see Notes below). Only in the former species are both tergite and sternite 8 much darker (Figs 106, 107) than in the latter one (Figs 108–110), but possibly this trait is variable, since in some cases there seem to be transitions in coloration.

Remarks. The attribution of these or those female

specimens to I. uniformis or the new species has been established, based of the following observations. Considering that I. uniformis, judging from the original description, is known from both parts of Malaysia and that I have at my disposal a male of I. uniformis and one

female (Fig. 8) from the same locality (W Malaysia, Perak, Cameron Highlands, Ringlet), both collected at about the same time (25.03–3.04.2002 and 26–31.03.2000, respectively), I have assigned them to the same species. Their both tergite and sternite 8 are coloured clearly light tones. Identically coloured are both tergite and sternite 8 in two females originating from Tanah Rata, located on the same, Cameron Highlands, as well as in most of the females from Eastern Malaysia, in particular, from the Crocker Range (Sabah), the type locality of I. uniformis, and from Trus Madi Mountain (Sabah). In the female collected in the latter locality (i.e. Trus Madi Mt.) on 23.03.2012 together with two males of the new species, including the holotype, as well as in one female from the same place (27.02.2014) and in another female from Pensiangan (Sabah), both tergite and sternite 8 are equally strongly darkened. I have preliminarily attributed them to paratypes of I. klimenkoi sp. n. By the character of the coloration of both tergite and sternite 8 in two females and one female originating from the Crocker Range and Trus Madi Mountain, respectively, it appears impossible to unequivocally assign them to either species. It is quite obvious that to securely identify females of both species, further detailed studies are needed.

Description. Body length 16.2–22.9 mm, humeral width 3.95–5.5 mm, thereby holotype 18.5 and 4.8 mm, respectively. Coloration of integument mainly combines reddish brown, dark reddish brown and red tones; eyes, partly mandibles, head dorsally or, besides this, partly pronotum black.

Head with well-developed antennal tubercles; with a sharp median groove between bases of antennae, growing even sharper between eyes and on vertex, as well as with a very sharp, deep, oblique groove behind inner part of each of dorsal lobes of eyes; genae moderately short; eyes well-developed, relatively strongly convex; submentum in basal part with a rough irregular sculpture, in apical part clearly or strongly impressed, smooth; neck with sharp transverse folds both ventrally and laterally; antennae of male one way or another reaching beyond apex of elytra by last antennomere, in female usually distinctly not reaching the apex of elytra, but sometimes can reach it; length ratio of antennomeres 1–11 in male (holotype taken as an example), 33 : 10 : 34 : 27 : 28 : 36 : 42 : 44 : 44 : 42 : 51, in female (one of the

paratypes taken as an example), 32 : 10 : 34 : 27 : 30 : 37 : 42 : 42 :

41 : 39 : 50; antennomere 1 with a heterogeneous, partly rough, rugose, dense puncturation; antennomere 2 subequal in length and width; antennomeres 3–5 of male slightly inflated in apical part, as in Fig. 10; apical inner angle of antennomeres 6–10 with a small, but well-expressed, sharp spine; last antennomere with a more or lees obvious appendage.

Pronotum distinctly longitudinal, 1.14–1.16 times as long as width, at base slightly or barely wider than at apex; with a sharp constriction near apex; on disc with coarse or rough tubercles and folds, at apex in middle part with a truncate, sometimes apically emarginate protrusion; near apex with a clear or very clear, usually more or less roundish, median tubercle.

Scutellum strongly narrowed towards apex, triangular, at the very apex usually narrowly rounded.

Elytra in male distinctly narrowed towards apex, in female about parallel-sided, 2.46–2.54 times as long as humeral width; with both a relatively rough and very small, double, very contrastingly differing puncturation characteristic of the genus; apical external angle broadly or obtusely rounded, sutural angle obtuse or narrowly rounded.

Prosternum with a very sharp, deep, transverse groove in front of middle, with heterogeneous, coarse or rough folds; prosternal


image


Figs 31–46. Imbrius Pascoe, 1866, pronotum and prosternum.

31–33, 39–40 – I. solodovnikovi sp. n.; 34 – I. similis Hüdepohl, 1990; 35–38, 41–42 – I. lineatus Pascoe, 1866; 43 – I. fedorenkoi sp. n.; 44–46 –

I. simulator Holzschuh, 2005. 31, 34, 39, 43 – holotypes; 32–33, 40, 44 – paratypes; 35–36, 41–42 – syntypes; 31–32, 34–35, 39, 41, 43–46 – males; 33, 36–38,

40, 42 – females.

Рис. 31–46. Imbrius Pascoe, 1866, переднеспинка и простернум.

31–33, 39–40 – I. solodovnikovi sp. n.; 34 – I. similis Hüdepohl, 1990; 35–38, 41–42 – I. lineatus Pascoe, 1866; 43 – I. fedorenkoi sp. n.; 44–46 –

I. simulator Holzschuh, 2005. 31, 34, 39, 43 – голотипы; 32–33, 40, 44 – паратипы; 35–36, 41–42 – синтипы; 31–32, 34–35, 39, 41, 43–46 – самцы;

33, 36–38, 40, 42 – самки.


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Figs 47–66. Imbrius Pascoe, 1866 and Mimimbrius gen. n., pronotum and puncturation fragment of basal part of elytra.

47 – I. ephebus Pascoe, 1866; 48–49 – I. diehli Hüdepohl, 1989; 50 – I. uniformis Holzschuh, 2010 (from Western Malaysia); 51 – I. allardi Hüdepohl, 1992; 52–53 – I. corrugatus Hüdepohl, 1990; 54 – I. acutipennis (Fisher, 1935); 55–57, 63 – M. subargenteus (Gressitt et Rondon, 1970), comb. n.; 58 –

M. dembickyi sp. n.; 59, 61 – M. micaceus (Pascoe, 1858), comb. n.; 60, 62 – M. geminatus (Holzschuh, 2005), comb. n.; 64 – I. lineatus Pascoe, 1866; 65 – I. solodovnikovi sp. n.; 66 – I. fedorenkoi sp. n. 47, 49, 51–52, 57–58, 65–66 – holotypes; 48, 53 – paratypes; 47–48, 50, 53, 59–62, 64 – females; 49, 51–52, 54–58, 63, 65–66 – males.

Рис. 47–66. Imbrius Pascoe, 1866 и Mimimbrius gen. n., переднеспинка и фрагмент пунктировки основной части надкрылий.

47 – I. ephebus Pascoe, 1866; 48–49 – I. diehli Hüdepohl, 1989; 50 – I. uniformis Holzschuh, 2010 (из Западной Малайзии); 51 – I. allardi Hüdepohl, 1992; 52–53 – I. corrugatus Hüdepohl, 1990; 54 – I. acutipennis (Fisher, 1935); 55–57, 63 – M. subargenteus (Gressitt et Rondon, 1970), comb. n.; 58 –

M. dembickyi sp. n.; 59, 61 – M. micaceus (Pascoe, 1858), comb. n.; 60, 62 – M. geminatus (Holzschuh, 2005), comb. n.; 64 – I. lineatus Pascoe, 1866; 65 – I. solodovnikovi sp. n.; 66 – I. fedorenkoi sp. n. 47, 49, 51–52, 57–58, 65–66 – голотипы; 48, 53 – паратипы; 47–48, 50, 53, 59–62, 64 – самки; 49, 51–52, 54–58, 63, 65–66 – самцы.


process with a very clear or strong apical tubercle; mesosternal process without tubercle dorsally, between coxae slightly or barely wider than prosternal process; mesosternum partly, metasternum and sternites with a small dense puncturation; metasternum with a clear, but not too sharp, median groove; last (visible) sternite at apex in male very broadly truncate, in female widely rounded; last (visible) tergite at apex widely rounded both in male and female.

Legs moderately short; femora claviform; tarsomere 1 very clearly shorter than tarsomeres 2 and 3 combined.

Recumbent setation on head dorsally, pronotum, scutellum, the very base of elytra red-yellow tones or, besides this, partly yellow, on antennae and legs combined with yellow and grey tones, on elytra, except for base, and on venter grey, on disc of pronotum with a pattern as in Figs 10, 11, on elytra continuous and uniform; head, pronotum, including a disc, the very base of elytra, prosternum, partly abdomen and legs, most of antennomeres in apical part and, besides this, on antennomeres 2–7 on inner side with more or less long, erect, partly suberect, sparse or individual, thin setae.

Genitalia as in Figs 98, 100, 102, 103, 106, 107.

Etymology. This new species is dedicated to the memory of late Alexey Alexandrovich Klimenko who untimely left us (1970–2017), a remarkable Russian traveller and connoisseur of Coleoptera. His magnificent collections and other material on some groups of longicorn beetles from the Oriental Region, especially from Borneo, which he generously shared with me, formed the basis for a number of important taxonomic findings and the discovery of new, sometimes surprising taxa described in this work.

Disribution. Eastern Malaysia.

Imbrius corrugatus Hüdepohl, 1990 (Figs 9, 12, 52, 53, 439, 440)

Imbrius corrugatus Hüdepohl, 1990a: 68. Type locality: Philippines, Luzon, Mountain Province (according to the original description and the label of the holotype).

Material. ♂, holotype, (ZSM) (Fig. 9), “Philippinen, Luzon, V.[19]86”,

Mountain Province”, “Holotypus ♂ Imbrius corrugatus mihi, Hüdepohl 1987” (Fig. 439); 1♀, paratype (ZSM) (Fig. 12), “Philippinen, Romblon, Sibuyan Is., Espana”, “Paratypus ♀ Imbrius corrugatus mihi, Hüdepohl 1987” (Fig. 440); 1♂, 1♀ (cAM), Philippines, N Luzon, Nueva Vizcaya, Santa Fe, 03.2014 (leg. N. Layron); 1♀ (cAM), Philippines, Leyte Island, Sogod, 05.2015 (leg. N. Layron).

Morphological notes. Body length 15.5–19.3 mm, humeral width 3.85–4.9 mm, thereby holotype male 15.8 and 3.9 mm, respectively (the body length of the holotype as indicated in the original description, namely, 13.5 mm, is erroneous).

Disribution. Philippines.

Imbrius acutipennis (Fisher, 1935) (Figs 22, 23, 54)

Dymasius acutipennis Fisher, 1935: 586. Type locality: Malaysia, Sabah, Kinabalu Mt., 5500 ft., Lumu Lumu (according to the original description and the label of the holotype).

Imbrius acutipennis: Hüdepohl, 1989a: 51; Heffern, 2013: 10; Lingafelter et al., 2014: 10, fig. 9c, d.

Material. ♀, holotype, by monotypy (USNM) (photograph); 1♂ (NHMD), E Malaysia, Sabah, Crocker Range, 04.2001 (local collector), “Imbrius acutipennis (Fisher), Ole Mehl det.”; 1♂ (cAM ex NHMD) (Fig. 22), same geographical label, “Imbrius acutipennis Fish., Ole Mehl det. 2010”; 2♂ (NHMD), same locality, 03.2003, “Imbrius acutipennis Fisher,

C. Holzschuh det. 2006, 2008”; 1♂, 1♀ (NHMD), E Malaysia, Sabah, Trus Madi Mt., 03.2004 (local collector), “Imbrius acutipennis (Fisher), Ole Mehl det. 2007”; 1♀ (NHMD), same locality, 04.2013, “Imbrius acutipennis (Fisher), Ole Mehl det. 2014”; 1♀ (cAN) (Fig. 23), Sabah, Trus Madi Mt.,

1160 m, 5.07.2011 (leg. A. Klimenko), “Imbrius ephebus Pascoe, 1866”; 1♀ (cAM), same geographical label; 5♀ (cAM), Sabah, Trus Madi Mt., 1250 m, 05°26’35”N / 116°27’5”E, 22–26.05.2014 (leg. A. Klimenko); 2♂ (cNO)

(photographs).

Morphological notes. Body length 18–24.3 mm, humeral width 3.9–5.45 mm.

Disribution. Eastern Malaysia.

Imbrius allardi Hüdepohl, 1992 (Figs 24, 51, 438)

Imbrius allardi Hüdepohl, 1992: 411. Type locality: [Malaysia] Sabah, Crocker Range (according to the original description and the label of the holotype). Heffern, 2013: 10.

Material. ♂, holotype (ZSM) (Fig. 24), “Borneo, Sabah, Crocker

Range, II.[19]90 [unknown collector]”, “Holotypus ♂ Imbrius allardi mihi, Hüdepohl 1991” (Fig. 438).

Morphological notes. Holotype male. Body length

24.6 mm, humeral width 6.25 mm.

Remarks. According to the personal communication of Mrs. Katja Neven (ZSM) of November 9, 2015, Hüdepohl’s collection contains only the holotype male which I have revised. However, based on the original description [Hüdepohl, 1992: 411], there must also be a paratype female. No other specimens of this species are known to me.

Disribution. Eastern Malaysia.

Key to species of Imbrius

  1. Apical sutural angle of elytra of one or another shape, often more or less straight, but in any case neither sharpened nor produced into a tooth 2

  2. Elytral setation of dense, recumbent, light setae forming more or less clear longitudinal strips, as in Figs 1–6, but in any case clearly neither uniform nor with dark v-shaped (or more or less similar) fascia in apical part formed by absence of dense light setae 3

  3. Longitudinal strips of elytra of dense, recumbent, light setae clearly narrower, as in Figs 1, 2, 4, 5; sculpture of pronotal disc noticeably coarser, as in Figs 31–33, 35–38; profemora near apex both on external and inner sides weakly or hardly impressed 4

  4. Disc of pronotum with a less strongly developed recumbent light setation, as in Figs 35–38; median longitudinal tubercle near apex of pronotum more strongly expressed, as in Figs 35–38; recumbent light setation of prosternum between constriction at its apex and anterior border of coxal cavities sparser, as in Figs 41, 42; apex of penis as in Figs 84, 85; parameres


    clearly narrower, as in Figs 89, 90, base of tegmen much narrower, as in Figs 89, 90 (W Malaysia, Singapore; Indonesia: Sumatra) .......................................... I. lineatus

  5. Setation of elytra of dense, recumbent, light setae entirely uniform, as in Figs 7–12 6

  6. Apical inner angle of antennomeres 6–10 with a clear or well-developed sharp spine 7

  7. Antennomeres 3–5 of male in apical part more strongly inflated, as in Fig. 7, antennomeres 3–7 or 3–8 of male on inner side with less numerous long erect and suberect setae; endophallus, apical part of penis and tegmen as in Figs 99, 101, 104, 105; both tergite and sternite 8 of female relatively light, as in Figs 108–110 (Indochina: W Malaysia; Borneo) ............... I. uniformis*

  8. Pronotum with a clearly or much more strongly

    developed recumbent light setation, as in Figs 47–49; antennae of male much longer than body, reaching beyond apex of elytra usually by antennomere 8, most of antennomeres much more elongated, as in Figs 13, 16, 18 9

  9. Antennomere 1, predominantly dorsally, with a quite coarse, very dense and confluent puncturation, creating a well-expressed scabrous surface; antennomere 3, 1.25–1.36 times as long as antennomere 4; body length 13.1–19.4 mm (rarely up to 19.8 mm); scutellar setation of recumbent dense setae usually clearly less; antennae of male slightly, but distinctly not reaching the base of elytra by antennomere 4 (Fig. 18) ................

    ............................................................................... I. ephebus

  10. Setation of pronotum on sides very weakly developed, as in Fig. 43; folds on disc of pronotum wider and more strongly rounded dorsally, not crest-shaped; setation of elytra with clearly creamy tone (Fig. 19); both tegmen and apical part of penis as in Figs 92, 95 (Indochina: S Vietnam) ........................................... I. fedorenkoi sp. n.

  11. Elytra strongly elongated, 2.9–3.1 times as long as humeral width, their apical sutural angle sharpened, as in Figs 22, 23, or drawn into a short tooth, with a more or less uniform, silky, mostly grey setation, sometimes appearing somewhat striped longitudinally; setation of pronotal disc of recumbent light setae well-developed, as in Figs 22, 23, 50; prosternal process without clear tubercle at apex; antennae of male reaching beyond apex of elytra by a penultimate antennomere (Fig. 22) ..

........................................................................ I. acutipennis

Genus Pachydissus Newman, 1838

Pachydissus Newman, 1838: 494. Thomson, 1864: 231;

Lacordaire, 1868: 265; Gemminger, 1872: 2804; Gahan, 1891:

24; 1906: 133; Aurivillius, 1912: 56; Gressitt, 1951: 141; Gressitt,

Rondon, 1970: 71; Adlbauer, 2002: 158; Catalogue..., 2010: 162;

Ślipiński, Escalona, 2016: 223.

Type species: Pachydissus sericus Newman, 1838, by monotypy.

Remarks. This genus is very widely distributed and covers Africa, South, East and Southeast Asia, Australia and Papua New Guinea, generally encompassing about 40 species [Ślipiński, Escalona, 2016; Base de données...,

2017].

Reviews of the African and Australian representatives of the genus were made by Adlbauer [2002] and Ślipiński and Escalona [2016], respectively. Papua New Guinea supports only two species [Montrouzier, 1855; Gressitt, 1959]. The Asian group of species needs a detailed revision and a diagnostic re-evaluation of the genus as a whole.

The first transformations in the supraspecific classification of the genus were implemented by Gahan


image

Figs 393–399. Falsopachydissus gen. n. and Pachydissus Newman, 1838, habitus, dorsal view, pronotum and last left antennomere.

393–397 – F. foveiscapus (Holzschuh, 2011), comb. n. (394 – paratype, after Holzschuh [2011], photograph by Luboš Dembický); 398 – P. parvicollis Gahan, 1891; 399 – P. schmutzenhoferi Holzschuh, 1990 (after Holzschuh [1990], photograph by Luboš Dembický). 398–399 – holotypes; 393, 397–399 – males; 394–396 – females.

Рис. 393–399. Falsopachydissus gen. n. и Pachydissus Newman, 1838, общий вид сверху, переднеспинка и последний левый членик усиков. 393–397 – F. foveiscapus (Holzschuh, 2011), comb. n. (394 – паратип, по [Holzschuh, 2011], фотография Л. Дембицкого); 398 – P. parvicollis

Gahan, 1891; 399 – P. schmutzenhoferi Holzschuh, 1990 (по [Holzschuh, 1990], фотография Л. Дембицкого). 398–399 – голотипы; 393, 397–399 – самцы;

394–396 – самки.


image

Figs 400–405. Pachydissus Newman, 1838 and Dymasius J. Thomson, 1864, habitus, dorsal view.

400 – P. sericus Newman, 1838; 401 – P. australasiae (Hope, 1842); 402 – P. intermedius Gahan, 1891; 403 – P. probatus Gahan, 1893; 404 – P. patricius Holzschuh, 1991 (after Holzschuh [1991], photograph by Luboš Dembický); 405 – D. querceus Holzschuh, 2015 (after Holzschuh [2015], photograph by Luboš Dembický). 400, 402–405 – holotypes; 400–402, 405 – males; 403–404 – females.

Рис. 400–405. Pachydissus Newman, 1838 и Dymasius J. Thomson, 1864, общий вид сверху.

400 – P. sericus Newman, 1838; 401 – P. australasiae (Hope, 1842); 402 – P. intermedius Gahan, 1891; 403 – P. probatus Gahan, 1893; 404 –

P. patricius Holzschuh, 1991 (по [Holzschuh, 1991], фотография Л. Дембицкого); 405 – D. querceus Holzschuh, 2015 (по [Holzschuh, 2015], фотография Л. Дембицкого). 400, 402–405 – holotypes; 400–402, 405 – самцы; 403–404 – самки.


[1891], who established four taxa as its subgenera (“sections”): Pachydissus s. str. (“verus”), Margites Gahan, 1891, Derolus Gahan, 1891 and Diorthus Gahan, 1891. However, he later [Gahan, 1906] promoted his taxa to full genera. This was accepted by Aurivillius [1912], Gressitt [1951], Gressitt and Rondon [1970] and subsequent researchers.

At the same time, Gressitt and Rondon [1970] proposed to divide Pachydissus into two subgenera, including Plavichydissus, described by Pic [1946] as a separate genus. But they noted thereby: “Plavichydissus... We are tentatively (sic) treating the latter as a subgenus of Pachydissus.” Until now, such a subgeneric structure is supported by some modern researchers (see, for example, Base de données... [2017]).

In the present work, it seems to me impossible to discuss in detail the systematic position of Plavichydissus and its morphological features. However, provisionally I would like to note that this taxon seems should not be considered within Pachydissus. Representatives of Plavichydissus are characterized at least by the peculiar sculpture and setation of the elytra and pronotum, as well as by the apex of the elytra devoid of clear or long teeth or spines, by non-inflated antennomeres 3 and 4 of the male, and only up to 28 mm in length.

When determining, again just preliminarily, the composition of Plavichydissus, besides its type species,

P. semiplicatus (Pic, 1926) [Pic, 1946], as well as Pachydissus (Plavichydissus) grossepunctatus Gressitt et Rondon, 1970, I would prefer to consider among its representatives, also included therein earlier [Pic, 1946], P. rufipennis (Pic, 1923) and such species as Margites sulcicollis Gahan, 1893,

M. decipiens Holzschuh, 1989, M. aggregatus Holzschuh, 1999 and M. sodalis Holzschuh, 1999. However, further studies are needed to clarify the diagnosis and species composition of the genus Plavichydissus.

Pachydissus foveiscapus was described relatively recently from Eastern Malaysia, but the author noted the following: “Differentialdiagnose. Die neue Art ist schwierig in eine der bekannten Gattungen unterzubringen. Nur wegen des ziemlich kurzen Halsschildes stelle ich sie vorläufig zu Pachydissus Newman, 1838.” [Holzschuh, 2011: 280–281]. Indeed, this species shows both a peculiar combination of features and some structural details that are generally typical of neither Pachydissus nor other similar genera. In this regard, it seems to me appropriate to distinguish P. foveiscapus in a separate, new genus described below.

Genus Falsopachydissus Miroshnikov, gen. n.

Type species: Pachydissus foveiscapus Holzschuh, 2011.

Diagnosis. This new monotypic genus, the single representative of which has hitherto been in the genus Pachydissus (see above), differs сlearly from the latter (except for one African species, keeping in mind only the structure of the antennae; see Remarks below) by the structure of the apical inner angle of antennomeres 5–10 with a long sharp spine, the last antennomere being strongly sharpened or extended apically into a very sharp

spine, as in Fig. 396, the coarser, partly one way or another shaggy, dense setation of the elytra, as in Figs 393–395, the somewhat peculiar distribution of the pronotum setation, as in Figs 393–395, 397, reminding of some Zatrephus.

Besides this, Falsopachydissus gen. n. is similar to neither Asian forms (see, for example, Figs 398, 399, 404) nor numerous other representatives of Pachydissus in a number of other features, differing by the more strongly convex pronotum on the disc, the shorter antennomere 1 with a strong or at least very clear impression near the base dorsally, the distinctly tuberous (more clearly so in the female) mesosternal process in the middle part on the sides, the shorter and wider submentum, the shape of the inflated

apical part of antennomeres 3 and 4 of the male (in contrast to species with thickened apices of male antennomeres 3 and 4), the more strongly elongated last antennomere of the male relative to the previous antennomere, the somewhat peculiar shape of the apical external angle of the elytra; differs also from some Asian species by the shorter legs with more robust tibiae, and the clearly shorter meta- and mesotarsi 1.

For the purpose of a more complete characterization of the new genus, it is also advisable to use some details of the structure of its sole representative as shown in the comparatively recent paper of Holzschuh [2011].

Very interesting is the strong similarity in the structure of the antennae of Falsopachydissus gen. n. to another new genus described above, Spinidymasius gen. n., but the former differs clearly from the latter by the more robust body; the wider head, including the broader frons and the more widely spaced bases of the antennae; the apical inner angle of antennomere 5 with a long sharp spine similar to the following antennomeres, except for the last one (in Spinidymasius gen. n., the apical inner angle of antennomere 5 can only be with a small denticle, but not a long spine); the structure of the pronotum, in particular, its being clearly more strongly convex on the disc and there bearing a somewhat peculiar sculpture and setation; the character of elytral setation (in Spinidymasius gen. n., elytra, if with a continuous setation, then clearly silky or distinctly patterned, but not shaggy).

At the same time, by the structure of the antennae, at least by the presence and location of the spine at an apical inner angle of antennomeres 5–10 and the shape of most antennomeres, the new genus is similar to some members of the genus Massicus Pascoe, 1867, in particular,

M. venustus (Pascoe, 1859), but differs very clearly in many features, including the more robust body, the much less strongly elongated elytra, the pattern of their recumbent setation, the structure of the pronotum, the stronger and shorter legs, etc.

Remarks. Among the African Pachydissus, only one species, P. samai Adlbauer, 2000, has the apical inner angle of the antennomeres 5–10 with a distinct spine, but in many other features it is a characteristic representative of the African group of species of this genus (Adlbauer [2002] and his personal communication of July 21, 2017). Besides this, in P. samai, in contrast to Falsopachydissus gen. n., the last antennomere is distinctly rounded apically (personal communication of Dr. Karl Adlbauer of July 21, 2017).


It is noteworthy that the apical inner angle of antennomeres 5–10 (in addition, sometimes the 3rd and 4th as well) carrying a more or less long spine, on the contrary, is characteristic of one or another representative of the genus Aeolesthes Gahan, 1890, with which some authors compare Pachydissus [Ślipiński, Escalona, 2016]. However, in Aeolesthes the last antennomere is the same as in Pachydissus, neither sharpened apically nor drawn into a sharp spine, but differs clearly from the new genus by the same features as Pachydissus does, including the setation of the pronotum and elytra, the sculpture of the pronotum and a number of other features.

Until now, I have been able to study more or less in due detail only the majority of Australian (see, for example, Figs 400–403), some Asian and several African representatives of Pachydissus (including the type species of the genus, the Australian P. sericus Newman, 1838, Fig. 400), while the structure of the other species from Australia, Asia and Africa, as well as the species from Papua New Guinea, have been studied, based on literature data alone [Gressitt, 1959; Holzschuh, 1991; Adlbauer, 2002; Ślipiński, Escalona, 2016 and others], as well as many quality photographs and personal advice kindly provided to me by Dr. Karl Adlbauer (Graz, Austria) on some African forms. Several species of this genus remain unknown to me, in particular, P. elegans Nonfried, 1895. Nevertheless, already at this stage I tend to believe that Pachydissus and Falsopachydissus gen. n. do not constitute a group closely related supraspecific taxa. At least in the future, in the case of a detailed revision of Pachydissus, the new genus can hardly be justified as one of the latter’s subgenera. No analogy with Dymasius and Spinidymasius gen. n. seems to be observed on this case (see above).

Etymology. In Latin, “falsus” means “false”, i.e., in general the name is treated as “false Pachydissus”.

Composition. The new genus includes a single species.

Disribution. Until now, this genus is known only from Borneo.

Falsopachydissus foveiscapus (Holzschuh, 2011),

comb. n.

(Figs 393–397)

Pachydissus foveiscapus Holzschuh, 2011: 279. Type locality: Malaysia, Sabah, Crocker Range (according to the original description). Heffern, 2013: 11.

Material. ♂, holotype (cCH) (photograph); 1♀, paratype (NHMD),

E Malaysia, Sabah, Trus Madi Mt., 03.2003 (local collector), “Paratypus Pachydissus foveiscapus n. sp., det. C. Holzschuh 2011”; 1♀ (cAM) (Fig. 395), same locality, 22–26.05.2014, 1250 m, 05°26’35”N / 116°27’5”E

(leg. A. Klimenko); 1♂ (NHMD), E Malaysia, Sabah, Crocker Range, 04.2014 (local collector), “Pachydissus foveiscapus, O. Mehl det. 2014”; 1♂ (BMNH) (Fig. 393), “Brunei, Kuala Belalong FSC [Field Studies Centre], 4.34°N, 115.7°E, Dipterocarp forest, Dryobalanops becarii, BM(NH), 1991–173”, “Aerial Malaise 3, 220 m alt., 8.VI.[19]91, N. Mawdsley [leg.] NM181”, “57”, “Zatrephus spinosus Brong., E. Vives det. 2005”, “Pachydissus” foveiscapus Holzschuh, 2011 ♂ det. A. Miroshnikov 2016”; 1♀, paratype (cDH) (photograph); 1♀, paratype (cCH) (photograph; Fig. 394).

Morphological notes. Body length 26–34 mm.

Disribution. This species has hitherto been known only from Eastern Malaysia [Holzschuh, 2011]. Based on the studied material, F. foveiscapus comb. n. is being recorded here from Brunei for the first time.

Genus Massicus Pascoe, 1867

Massicus Pascoe, 1867: 319 [name replacement for Conothorax J. Thomson, 1864, non Jeckel, 1854 (Curculionidae)]. Lacordaire, 1868: 260; Gemminger, 1872: 2802; Gahan, 1906: 129;

Aurivillius, 1912: 55; Gressitt, Rondon, 1970: 59; Lee, 1982: 27;

Kusama, Takakuwa, 1984: 252; Hüdepohl, 1990a: 60; Catalogue...,

2010: 161; Heffern, 2013: 10.

Mallambyx Bates, 1873: 152 (Pachydissus subgen.).

Mallambyx: Ganglbauer, 1889: 473; Winkler, 1929: 1141;

Matsushita, 1933: 243; Mitono, 1940: 82; Plavilstshikov, 1940: 79,

635; Gressitt, 1951: 135; Kojima, Hayashi, 1969: 47; Tsherepanov,

1981: 22.

Type species: Cerambyx pascoei J. Thomson, 1857, by monotypy.

Remarks. This genus is widely distributed and covers South, East and Southeast Asia. Taking into account the new forms described here, it contains no less than 15 species.

Massicus is very similar to Neocerambyx J. Thomson, 1861, but the diagnoses of both genera require a further detailed development, since the morphological differences between them as proposed by various researchers are generally unstable and can be used only for part of the species. Neocerambyx or Massicus (= Mallambyx Bates, 1873) raddei Blessig, 1872 can be mentioned as a striking example, when many publications treat the same species in different genera [Blessig, 1872; Heyden, 1880–1881; Ganglbauer, 1889; Pic, 1900; Aurivillius, 1912; Winkler,

1929; Matsushita, 1933; Mitono, 1940; Plavilstshikov, 1940;

Gressitt, 1951; Kojima, Hayashi, 1969; Tsherepanov, 1981; Lee, 1982; Kusama, Takakuwa, 1984; Hüdepohl, 1990b; Hua, 2002; Catalogue..., 2010 and many others].

Descriptions of two new species from Malaysia are given below.

Massicus valentinae Miroshnikov, sp. n.

(Figs 406, 409)

Material. Holotype, ♂ (cAM) (Fig. 406): W Malaysia, Pahang, Cameron Highlands, Tanah Rata, 04.2015 (local collector). Paratype: 1♀ (cAM), W Malaysia, Pahang, Bukit Fraser [= Fraserʼs Hill], 3°43’N / 101°44’E, 1000–1300 m, 9–11.05.2007 (leg. V. Tuzov).

Diagnosis. The new species seems to be especially similar to M. trilineatus (Pic, 1933) and M. taiwanus Makihara et Niisato, 2014, but differs from both by the clearly more strongly convex pronotum; the somewhat peculiar, recumbent, light setation on its disc forming clearly narrower longitudinal strips and significantly masking the sculpture in the middle part of the disc (especially so in the male), due to which the longitudinal strips look less contrasting against the general background of the setation, as in Fig. 409 (cf. Figs 412, 413); the shorter and more obtuse tooth of the sutural angle of the elytra; the structure of the apical external angle of antennomeres 5–10 or 6–10, each bearing a well-developed, sharp spine; the somewhat shorter antennae of the male; antennomeres 3 and 4 slightly more strongly inflated apically in the male. Massicus valentinae sp. n. can also be compared to M. pascoei (J. Thomson, 1857), but differs clearly by the structure of the pronotum, including its shape and setation, as in Fig. 409 (cf. Fig. 414); the slenderer antennae; the smoothened sculpture and the less strongly protruding apical external


image

Figs 406–414. Massicus Pascoe, 1867, habitus, dorsal view, and pronotum.

406, 409 – M. valentinae sp. n.; 407, 408, 410–411 – M. ivani sp. n.; 412 – M. trilineatus (Pic, 1933); 413 – M. taiwanus Makihara et Niisato, 2014;

414 – M. pascoei (J. Thomson, 1857), male. 406–407, 409–410 – holotypes, males; 408, 411 – paratype, female.

Рис. 406–414. Massicus Pascoe, 1867, общий вид сверху и переднеспинка.

406, 409 – M. valentinae sp. n.; 407, 408, 410–411 – M. ivani sp. n.; 412 – M. trilineatus (Pic, 1933); 413 – M. taiwanus Makihara et Niisato, 2014;

414 – M. pascoei (J. Thomson, 1857), самец. 406–407, 409–410 – голотипы, самцы; 408, 411 – паратип, самка.


angle of antennomere 1; the much less strongly expressed individual punctures (discarding very small puncturation) on the 3rd and some following antennomeres; the more strongly developed spine on antennomeres 5–10 or 6–10; the slightly shorter antennae in the male; the shape of the inflated apical part of antennomeres 3 and 4 in the male; the shape of the apical external angle of the elytra; the clearly longitudinal antennal tubercles; the structure of the submentum; the seemingly more uniformly curved external margin of the mandibles; the slightly more strongly convex eyes; and on the average a smaller body. The differences between M. valentinae sp. n. and another similar new species, M. ivani sp. n., are shown below in the diagnosis of the latter.

Description. Body length 41.5 or 52.4 mm, humeral width

9.8 or 13 mm in male and female, respectively. Head, pronotum, basal antennomeres, prosternum, except for process partly, mesosternum, tibiae and tarsi entirely or almost completely black; elytra, apical antennomeres, prosternal process partly, metasternum, sternites, femora, most of coxae red-brown and dark red-brown tones.

Head with moderately developed antennal tubercles; with a distinct, narrow, median groove between bases of antennae, a wider and deep one between eyes and partly on vertex; with a dense, partly sharp puncturation dorsally; eyes moderately convex; longitudinal diameter of lower lobe of eye 1.28 or 1.5 times as long as genae in female and male, respectively; submentum with a rough, partly obliterated puncturation, in middle part with a well-expressed transverse fold, in front of and behind which with clear depressions; neck almost entirely with more or less sharp transverse folds ventrally; gula also with sharp transverse folds; antennae of male significantly longer than body, reaching beyond apex of elytra by basal part of antennomere 8, in female reaching beyond apex of elytra by penultimate antennomere; length ratio of antennomeres 1–11 in male, 38 : 8 : 67 : 42 : 61 : 78 : 77 : 70 :

59 : 56 : 96, in female, 46 : 10 : 77 : 44 : 60 : 71 : 67 : 58 : 52 : 49 : 66;

antennomere 1 without coarse sculpture, only with a relatively small dense puncturation, apical external angle very clearly protruding and obtusangularly; antennomere 2 distinctly transverse; antennomere 3 clearly, antennomeres 4 and 5 less strongly inflated at apex; apical external angle of antennomeres 5–10 or 6–10 with a sharp spine, more or less equally developed on all these antennomeres; last antennomere sharpened apically.

Pronotum 1.02 or 1.05 times as wide as long, base 1.29 or

1.41 times as wide as apex in male and female, respectively; relatively strongly (male) or very clearly (female) convex; on disc with a very coarse, transverse, partly irregular folds, as in Fig. 409.

Scutellum strongly narrowed towards apex, triangular.

Elytra 2.64–2.8 times as long as humeral width; moderately convex; with a small dense puncturation; apical external angle rounded, sutural angle with a short, but distinct tooth almost entirely masked under a dense setation.

Prosternum in apical one-third at middle visibly tuberculiform elevated, mainly with transverse coarse folds in apical part; prosternal process distinctly broadened towards apex dorsally, with a clear or very clearl tubercle at apex; mesosternal process without tubercle dorsally, between coxae clearly wider than prosternal process; meso- and metasterna and sternites with a small dense puncturation; metasternum with a sharp median groove; both last (visible) sternite and tergite of male truncate apically; last (visible) sternite in female truncate, last (visible) tergite with a weak emargination.

Legs moderately long; tarsomere 1 barely shorter than tarsomeres 2 and 3 combined.

Recumbent dense setation ocher-yellow, entirely or almost completely clothing head dorsally, elytra, scutellum, antennomeres 1 and 2, pro-, meso- and metasterna, sternites, legs,

mandibles dorsally, partly antennomere 3; setation of pronotum as in Fig. 409 (see also Diagnosis above); head, pronotum on disc and laterally, antennomeres 3–10 in apical part, apex of abdomen, legs mainly on trochanters with more or less long, erect, partly suberect, sparse or individual, thin setae.

Etymology. I am very pleased to dedicate this new species to my mother, Valentina Dmitrievna Miroshnikova, who celebrated her 90th birthday on September 29, 2017 and whose constant care for my family I feel all the time.

Disribution. Western Malaysia.

Massicus ivani Miroshnikov, sp. n.

(Figs 407, 408, 410, 411)

? Massicus trilineatus trilineatus auct.: Heffern, 2005: 16 (partim, Borneo) (non Pic, 1933); Heffern, 2013: 10 (partim, Borneo) (non Pic, 1933).

Material. Holotype, ♂ (NHMD) (Fig. 407): E Malaysia, Sabah,

Keningau Distr., Trus Madi Mt., 03.2005 (local collector). Paratype: 1♀ (cAM ex NHMD) (Fig. 408), same label as holotype.

Diagnosis. The new species seems to be especially similar to M. valentinae sp. n., but differs by the somewhat more robust body and antennae, especially so the basal antennomeres; the shorter antennomere 3; the apical external angle of antennomere 6 with a less strongly developed denticle, but more robust (not so long) one on the following antennomeres, except for the last one; the structure of the pronotum, in particular, it being less strongly convex on the disc at least in the male and somewhat less strongly broadened at the middle in the male; the much less strongly developed recumbent light setation in the middle part of the disc only weakly masking its sculpture there, as in Figs 410, 411 (cf. Fig. 409); the apical sutural angle of the elytra of the male with a more strongly developed tooth; the sharper tubercle at the apex of the prosternal process; the somewhat more strongly elongated mesosternal process of the male; the more robust legs, at least so in the male, as in Fig. 407 (cf. Fig. 406); and the larger body in the male and female, respectively. Massicus ivani sp. n. can also be compared to M. pascoei, but differs, partly like

M. valentinae sp. n., in the structure of the antennae, in

particular, the smoothened sculpture and the less strongly protruding apical external angle of antennomere 1; the more strongly developed denticle on antennomeres 7–10; many antennomeres less strongly elongated, including 3rd and 4th, in which, additionally, the apical part is somewhat differently inflated in the male; the shorter antennae in both sexes; the somewhat peculiar sculpture of the pronotum, as in Figs 410, 411 (cf. Fig. 414); the less strongly developed sutural tooth at the apex of the elytra; the clearly rounded apical external angle of the elytra; the shape of the antennal tubercles and the external margin of the mandibles; the seemingly slightly more strongly convex eyes; the shape and sculpture of the submentum and some other features.

Besides this, by the habitus, larger body, partly the

structure of the antennae and some other traits the new species resembles Neocerambyx pubescens Fisher, 1936 (see Note), but differs in the slenderer basal antennomeres; antennomere 1 clearly less strongly broadened towards the apex, with a smooth sculpture; the apical external angle of antennomeres 7–10 with a more strongly developed denticle; the more closely spaced dorsal lobes of the eyes;


the significantly longer median groove on the vertex, mainly hidden by dense setation between the bases of the antennae; the less transverse pronotum and its generally less strongly developed setation; the sculpture on the ventral side of the head; and the more strongly elongated and narrower process of the mesosternum in the male.

At the same time, evident similarities of M. ivani sp. n. and M. unicolor Gahan, 1906 must be noted in the structure of the antennae, the shape of the elytral apex and some other details, but the former species differs clearly from the latter at least by the coloration of the recumbent setation of the dorsum, the less strongly protruding apical external angle of antennomere 1, the structure of the pronotum, including its somewhat peculiar shape and sculpture, and the distribution of recumbent setation, the shorter

antennae of the female in which many antennomeres are less strongly elongated.

Note. I have only been able to examine the holotype male of Neocerambyx pubescens, kept in the USNM, from a series of high-quality photographs of the habitus and some structural details, all kindly provided by Dr. Alexandr S. Konstantinov (USNM) upon my request.

Until recently N. pubescens was only known from Java. Based on the data available at the Insecterra Forum (http://insecterra.forumactif.com/), this species has also been recorded from Borneo, at the same locality as

M. ivani sp. n. (Sabah, Trus Madi Mt.: http://insecterra. forumactif.com/t24742-massicus).

Description. Body length 56.2 or 65 mm, humeral width 13.6 or 16.8 mm in male and female, respectively.

Head, pronotum, antennae, prosternum, except for process partly, mesosternum, tibiae and tarsi entirely or almost completely black; elytra, prosternal process partly, metasternum, sternites, femora, predominantly coxae red-brown and dark red-brown tones; elytra in male with a more strongly expressed red tint than that in female.

Head with very strong antennal tubercles; with a distinct, narrow, median groove between bases of antennae, as well as with a wider one between eyes and partly on vertex, a well-expressed, but moderately deep in male and a significantly deeper in female; with a dense puncturation dorsally, coarser in female; eyes moderately convex; genae slightly shorter than longitudinal diameter of lower lobe of eye; submentum with puncturation rough, predominantly obliterated in male, clearer in female, in middle part with a coarse transverse fold roundish dorsally in male, in front of and behind which with well-expressed depressions; neck ventrally in middle part almost smooth, lateral to middle with more or less sharp transverse folds; gula with transverse folds, very sharp in male, weak in female; antennae of male much longer than body, reaching beyond apex of elytra by antennomere 8, in female noticeably not reaching the apex of elytra; length ratio of antennomeres 1–11 in male, 31 : 8 : 43 : 31 : 45 : 55 : 58 : 56 : 50 : 46 : 80, in female, 31 : 8 :

38 : 26 : 34 : 37 : 37 : 31 : 29 : 26 : 35; antennomere 1 without

coarse sculpture, only with a relatively small dense puncturation, apical external angle very clearly protruding and obtusangularly; antennomere 2 distinctly transverse; in male, antennomeres 3 and 4 pretty robust, antennomere 3 clearly, 4th and 5th less strongly inflated at apex; apical external angle of antennomeres 6–10 with a sharp denticle, most developed and sharper on antennomeres 8–10; last antennomere sharpened apically, in female by a shorter cone.

Pronotum 1.03 or 1.15 times as wide as long, at base 1.21 or 1.49 times as wide as apex in male and female, respectively; moderately convex in male and somewhat more strongly so in female; on disc with a very coarse, transverse, partly sinuous, in male partly irregular folds, as in Figs 410, 411.

Scutellum in male strongly narrowed towards apex, triangular, in female almost uniformly rounded towards apex, starting from base.

Elytra 2.6–2.66 times as long as humeral width; very clearly convex; with a small dense puncturation; apical external angle rounded; sutural angle in male drawn into a relatively long, well- visible tooth, in female with a barely expressed denticle masked under a dense setation.

Prosternum in apical one-third at middle clearly tuberculiform elevated, with transverse, partly coarse and oblique folds in apical part; prosternal process very clearly broadened towards apex dorsally, with a strong tubercle at apex, wider in female; mesosternal process without tubercle dorsally, between coxae clearly wider than prosternal process; meso- and metasterna and sternites with a small dense puncturation; metasternum with a sharp median groove; last (visible) sternite at apex in male broadly slightly rounded, almost truncate, in female with a narrow shallow emargination; last (visible) tergite at apex in male distinctly broadly rounded, in female very clearly obtusangularly emarginate.

Legs moderately long; tarsomere 1 barely or slightly shorter than tarsomeres 2 and 3 combined.

Recumbent dense setation ocher-yellow, entirely or almost completely clothing elytra, antennomeres 1 and 2, sternites, legs, most of head dorsally, pro- and mesosterna, partly scutellum in male, partially antennomere 3; setation of pronotum as in Figs 410, 411 (see also Diagnosis above); head, pronotum on disc and laterally, most of antennomeres in apical part, apex of abdomen, legs mainly on trochanters with more or less long, erect, partly suberect, sparse or individual, thin setae.

Etymology. This new species is dedicated to the memory of my father, Ivan Polikarpovich Miroshnikov (1928–1995), whose wise advice and very strong support I received during all his life.

Distribution. Eastern Malaysia.

Genus Sebasmia Pascoe, 1859

Sebasmia Pascoe, 1859: 18. Thomson, 1864: 234; Lacordaire,

1868: 272; Gemminger, 1872: 2807; Gahan, 1906: 143; Aurivillius,

1912: 61; Heffern, 2013: 11.

Type species: Sebasmia templetoni Pascoe, 1859, by monotypy.

Remarks. This Oriental genus includes eight species, five of them described relatively recently, while one species is described as new. One species previously referred to

this genus [Gahan, 1906: Sebasmia nigra] has since been transferred to Microdymasius Pic, 1946 [Holzschuh, 2015].

Sebasmia is characterized by a peculiar combination of features [Holzschuh, 2005] and differs clearly from all similar genera. Only this genus seems to show one very notable character distinguishing it from any other in the tribe. In the male, the middle part or most of the ventral side of the mesofemur has a dense or very dense brush consisting of coarse, sometimes partly twisted, light setae which can be located in a more or less clear or even deep emargination.

Representatives of the genus have hitherto been known only from Sri Lanka, Borneo and Sumatra [Pascoe, 1859; Gahan, 1906; Aurivillius, 1912; Heffern, 2005, 2013;

Holzschuh, 2005, 2006; Makihara et al., 2008; Base de

données..., 2017].

Based on the material studied (see below), the genus Sebasmia is being recorded here from Indochina for the first time.


image

Figs 415–420. Sebasmia Pascoe, 1859, habitus, dorsal view.

415–416 – S. speculifera Holzschuh, 2005 (415 – after Holzschuh [2005], photograph by Luboš Dembický); 417–419 – S. indochinensis sp. n. (417, 419 – from Vietnam, 418 – from Thailand); 420 – S. vetusta Holzschuh, 2006 (from Western Malaysia). 415, 417 – holotypes; 416, 418–419 – paratypes;

415–418, 420 – males; 419 – female.

Рис. 415–420. Sebasmia Pascoe, 1859, общий вид сверху.

415–416 – S. speculifera Holzschuh, 2005 (415 – по [Holzschuh, 2005], фотография Л. Дембицкого); 417–419 – S. indochinensis sp. n. (417, 419 –

из Вьетнама, 418 – из Таиланда); 420 – S. vetusta Holzschuh, 2006 (из Западной Малайзии). 415, 417 – голотипы; 416, 418–419 – паратипы; 415–418,

420 – самцы; 419 – самка.


image

Figs 421–434. Sebasmia Pascoe, 1859, details of structure.

421, 425, 429–430 – S. speculifera Holzschuh, 2005 (429 – photograph by Luboš Dembický); 422–424, 426–428, 431–434 – S. indochinensis sp. n.

(422–423, 426–427, 431–432, 434 – from Vietnam, 424, 428, 433 – from Thailand). 422, 426, 429, 431 – holotypes; 421, 423–425, 427–428, 430, 432–434 –

paratypes; 425–433 – males; 434 – female; 421–424 – apical part of tegmen, ventral view; 425–428 – apical part of male tergite 8, dorsal view; 429–434 – head, dorsal view, and pronotum.

Рис. 421–434. Sebasmia Pascoe, 1859, детали строения.

421, 425, 429–430 – S. speculifera Holzschuh, 2005 (429 – фотография Л. Дембицкого); 422–424, 426–428, 431–434 – S. indochinensis sp. n.

(422–423, 426–427, 431–432, 434 – из Вьетнама, 424, 428, 433 – из Таиланда). 422, 426, 429, 431 – голотипы; 421, 423–425, 427–428, 430, 432–434 –

паратипы; 425–433 – самцы; 434 – самка; 421–424 – вершинная часть тегмена снизу; 425–428 – вершинная часть 8-го тергита сверху; 429–434 – голова сверху и переднеспинка.


Sebasmia indochinensis Miroshnikov, sp. n.

(Figs 417–419, 422–424, 426–428, 431–434)

Material. Holotype, ♂ (BMNH ex cTT) (Fig. 417): Vietnam, Kon Tum Prov., Ngọc Linh, 1700 m, 04.2016 (local collector). Paratypes: 1♂ (cAM ex cTT), Vietnam, Thua Thien-Hue Prov., Bach Ma Mt., 1400 m, 16°11’N / 107°51’E, 03.2016 (local collector); 1♀ (cTT), same locality, 06.2017 (local collector); 1♀ (cAM) (Fig. 419), Vietnam, Lam Dong Prov., 25 km NNW of Bao Loc, Loc Bao env., 800 m, 11°44’18”N / 107°42’08”E, 5–20.04.2013 (leg. D. Fedorenko); 1♀ (cTT), Vietnam, Quang Nam Prov., Tay Giang Distr., Axan, 1300 m, 06.2017 (local collector); 1♂ (cTT), same locality, 07.2017 (local collector); 1♀ (cFV), Vietnam, Quang Nam Prov., 05.2017 (leg. N. Tai); 1♀ (cWT), Vietnam, Da Nang, BaNa Mt., 1450 m, 05.2014 (local collector); 1♀ (cTT), same locality, 05.2015 (local collector); 1♂ (cSM) (Fig. 418), N Thailand, Chiang Dao Hill Resort, 100 km N Chiang Mai, 600 m, 10–23.03.2010 (leg. S. Murzin).

Comparative material. Sebasmia speculifera Holzschuh,

2005: ♂, holotype (cCH) (photograph; Fig. 415); 1♂, paratype (cDH) (Fig. 416), Malaysia, Sabah, Trus Madi Mt., 17.03.2000 (local collector), “Cerambycini n. gen., n. sp. det. K. Hüdepohl 2001”, “Paratypus Sebasmia speculifera n. sp. det. C. Holzschuh 2004”.

Diagnosis. This new species is very similar to

S. speculifera, but differs at least by the clearly less strongly developed recumbent light setation both of the pronotum and head dorsally, especially of the former, as in Figs 431–434 (cf. Figs 429–430), the male genitalia, including the much shorter parameres, as in Figs 422–

424 (cf. Fig. 421), the somewhat peculiar shape of the apex of tergite 8, as in Figs 426–428 (cf. Fig. 425); on the average a larger body, as well as by the variable composition of the coloration of the pronotal setation, up to the complete absence of yellow/yellowish tones and the presence of only a silver-white tone, as in Fig. 432, in most cases the slightly longer male antennae, as in Figs 417–418 (cf. Figs 415, 416), and some other minor traits.

Description. Body length 17.9–21.7 or 24–26.4 mm, humeral width 3.4–4.2 or 4.7–5.2 mm in males and females, respectively, thereby holotype largest among the males, while Thai male smallest (body length of type specimens of S. speculifera, including of two female, 13.6–19.2 mm). Coloration of integument mainly combines black and dark reddish brown tones; eyes black; antennae and legs red-brown, antennomere 1 darkest, but femora can also be darker.

Head with well-developed antennal tubercles; with a sharp median groove between bases of antennae and a more or less sharp, sometimes weak groove between eyes partly and on vertex; genae relatively short; eyes of male very large and very strong convex, in female less strongly developed and less strongly convex, in both sexes with large ocelli; submentum with coarse punctures; neck with sharp transverse folds both ventrally and laterally; antennae of male reaching beyond apex of elytra by penultimate antennomere, sometimes by basal part of last antennomere, in female distinctly not reaching the apex of elytra or reaching it; length ratio of antennomeres 1–11 in male (holotype taken as an example) 32 : 11 : 37 : 35 : 52 : 58 : 60 : 58 : 57 : 55 : 68, in female (one of the paratypes taken as an example) 34 : 12 : 44 : 33 : 48 : 50 : 48 : 46 : 45 : 44 : 61; antennomere 1

mostly with a coarse, very dense and partly confluent, or mainly confluent puncturation; antennomere 2 barely longitudinal (given its longest inner side); last antennomere with a distinct appendage.

Pronotum clearly/very clearly longitudinal, 1.2–1.29 times as long as width; base 1.14–1.18 times as wide as apex; broadly rounded on sides; with a sharp or very sharp constriction near apex; with coarse or very coarse, mainly irregular, partly sinuous folds and with a strong or very strong, median, keel-shaped,

longitudinal, shiny dorsally elevation located mostly in basal half, as in Figs 431–434.

Scutellum triangular, at apex more or less narrowly truncate. Elytra strongly elongated, in male barely narrowed towards apex, in female about parallel-sided, 3.14–3.45 times as long as humeral width; slightly obliquely and broadly truncate at apex; sutural angle with a clear or very clear tooth; with somewhat heterogeneous, small, mostly gentle, sometimes sharper

puncturation.

Prosternum with rough or coarse, irregular, partly transverse folds in apical part and with a transverse, more or less clearly expressed groove in front of middle; prosternal process with a well-developed, sometimes weakly expressed, apical tubercle; mesosternal process without tubercle dorsally; metasternum and sternites with a clear, dense, small puncturation, somewhat heterogeneous in former; last (visible) sternite at apex in both sexes with a small or more strong developed emargination.

Legs relatively short; femora with a carina along each side; mesofemora in male with a well-developed brush of very dense light setae characteristic of the genus; tarsomere 1 slightly shorter than tarsomeres 2 and 3 combined.

Recumbent setation on head dorsally mostly or completely yellow/yellowish, as in Figs 431–434; on pronotum combined with yellow/yellowish and silver-white tones or entirely silver- white, thereby forming near apex no narrow fascia (Figs 431–434) (in S. speculifera, along with a clearly more strongly developed setation than in S. indochinensis sp. n., a very well-expressed, more or less narrow fascia located near apex, as in Figs 429, 430); on elytra forming longitudinal strips of silver-white or, in addition, partly yellowish tones, thereby in front of apex creating a relatively large naked spot of triangular or oval shape on each elytron, as in Figs 417–419; on venter completely or almost entirely silver-white; elytral apex strongly or entirely masked under a dense setation; head, pronotum laterally, prosternum partly, legs on trochanters with more or less long, erect, partly suberect, sparse or individual, thin setae.

Genitalia of male as in Figs 422–424, 426–428; parameres very short, peculiar in shape; tergite 8 truncate apically, thereby can be slightly emarginate (while in S. speculifera, parameres moderately long, tergite 8 broadly rounded apically, as in Figs 421, 425).

Etymology. The formation of the name of this new species is related to its distribution in some countries of Indochina Peninsula.

Distribution. Vietnam, Thailand.

Sebasmia vetusta Holzschuh, 2006 (Fig. 420)

Sebasmia vetusta Holzschuh, 2006: 228. Type locality: Malaysia, Sabah, Crocker Range (according to the original description). Heffern, 2013: 11 (Borneo).

Material. ♂, holotype (cCH) (photograph); 1♂ (BMNH) (Fig. 420),

“[Malaysia] Penang Hill, 2.260, Penang, 98–89”, “Sebasmiavetusta Holzschuh, 2006 ♂ det. A. Miroshnikov 2016”; 1♂ (cLD), Indonesia, Kalimantan Selatan, 40 km E Kandangan, Loksado, 2°30’S / 115°20’E, 7–22.09.1997 (leg. S. Jakl), “Sebasmia vetusta Holzschuh, 2006 ♂ det. A. Miroshnikov 2017”; 1♀ (cAM), E Malaysia, Sabah, Trus Madi Mt., 1160 m, 24.08.2012 (leg. A. Klimenko); 1♂ (cAM), E Malaysia, Sabah, Nabawan Distr., ~7 km N Pensiangan, 530 m, 04°35’16”N / 116°19’27”E, 29.05.2014 (leg. A. Klimenko); 1♀, paratype (cDH) (photograph).

Remarks. This species has hitherto been known only from Eastern Malaysia [Holzschuh, 2006]. Sebasmia vetusta is being recorded here from Western Malaysia and generally from Indochina, on the one hand, and from Indonesia (south of Borneo), on the other hand, for the first time.


image

Figs 435–461. Cerambycini Latreille, 1802, labels of types and other specimens.

435–436 – syntypes; 437–439, 441–442, 444–449, 451–453 (452 – photograph by Gérard Tavakilian), 456, 458–461 – holotypes; 440, 443 – paratypes; 454 – lectotype; 457 – “holotypus” (see Remarks to Zatrephus lumawigi Hüdepohl, 1990).

Рис. 435–461. Cerambycini Latreille, 1802, этикетки типовых и других экземпляров.

435–436 – синтипы; 437–439, 441–442, 444–449, 451–453 (452 – фотография Ж. Тавакиляна), 456, 458–461 – голотипы; 440, 443 – паратипы; 454 – лектотип; 457 – «holotypus» (см. замечания к Zatrephus lumawigi Hüdepohl, 1990).


Acknowledgements

I am very grateful to Svetlana V. Andreeva and Andrey M. Shapovalov (ZIN), Michael Balke, Katja Neven and Ditta Amran-Balke (ZSM), Maxwell V.L. Barclay and Michael F. Geiser (BMNH), James H. Boone (BM), Thierry Deuve, Azadeh Taghavian and Gérard L. Tavakilian (MNHN), Alain Drumont (IRSN), AlexandrS.Konstantinov(USNM),KirillV.Makarov(PUM), Alexey Yu. Solodovnikov and Sree Gayathree Selvantharan (NHMD) for the opportunity to study the museum material, to Gérard Chemin (Champigny-sur-Marne, France), Luboš Dembický (Brno, Czech Republic), Pierre Haller (Essert, Le Mouret, Switzerland), Daniel J. Heffern (Houston, USA), Philippe Jacquot (Montboucher-sur- Jabron, France), Sergey V. Murzin (Moscow, Russia), Alexandr Napolov (Riga, Latvia), Tomáš Tichý (Opava, Czech Republic), Francesco Vitali (Luxembourg), William H. Tyson (Coarsegold, USA) who have provided various specimens from their private collection. I would like to express my sincere thanks to Alexey Yu. Solodovnikov for his kind permission to retain some material in my personal collection, to Denis G. Kasatkin (Rostov- on-Don, Russia) for having rendered his great help in performing and studying the endophallus preparations, to Dmitry N. Fedorenko (Institute for Problems of Ecology and Evolution, Moscow, Russia), who was funded by the Russia-Vietnam Tropical Center, for the valuable material he rendered to me for study. I am deeply indebted to Alexandr G. Kirejtshuk (ZIN), again to Sergey V. Murzin, Alexey Yu. Solodovnikov who helped a lot in my prompt receipt of the material for study, to Karl Adlbauer (Graz, Austria), Sergey A. Kurbatov (Moscow, Russia), Nobuo Ohbayashi (Kamimiyada, Miura City, Japan), Andre Skale (Hof, Germany), Eduard Vives (Barcelona, Spain), again to Daniel J. Heffern, Philippe Jacquot, Alexandr G. Kirejtshuk, Alexandr S. Konstantinov, Kirill V. Makarov, Katja Neven, Sree Gayathree Selvantharan, Alexey Yu. Solodovnikov, Gérard L. Tavakilian, Tomáš Tichý, Francesco Vitali for the helpful provision of various pictures and/or valuable information, to Dmitry Telnov (Entomological Society of Latvia, Riga) for his efforts to help me locate some of the relevant material in several repositories. I give special thanks to Kirill V. Makarov for having rendered his great help in the preparation of almost all photographs, and to Luboš Dembický who generously shared the pictures of the holotypes of many species of the tribe Cerambycini. Last but not least, I am most grateful to my wife Tatiana who helped a lot in the preparation of the illustrations for publication.

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Received / Поступила: 2.11.2017 Accepted / Принята: 27.11.2017