_9_Lillig/Image_001.jpg)
Кавказский энтомол. бюллетень 12(2): 253–254 © CAUCASIAN ENTOMOLOGICAL BULL. 2016
Bund für Umwelt und Naturschutz Deutschland, LV Saarland, Evangelisch-Kirch-Straße 8, Saarbrücken 66111 Germany. E-mail: martin.lillig@t-online.de
Key words: Coleoptera, Tenebrionidae, Hypophloeus japonicus, Tribolium castaneum, new synonymy.
Ключевые слова: Coleoptera, Tenebrionidae, Hypophloeus japonicus, Tribolium castaneum, новая синонимия.
Abstract. The unique type specimen of the hitherto overlooked Japanese tenebrionid beetle Hypophloeus japonicus Thunberg, 1822 was studied and designated as lectotype. The new synonymy is established: Tribolium castaneum (Herbst, 1797) = Hypophlaeus japonicus Thunberg, 1822, syn. n.
Резюме. Типовой экземпляр до настоящего времени забытого вида Hypophloeus japonicus Thunberg, 1822 был изучен и обозначен как лектотип. Устанавливается новая синонимия: Tribolium castaneum (Herbst, 1797) = Hypophlaeus japonicus Thunberg, 1822, syn. n.
During the preparation of a world catalogue of the Hypophlaeini (Lillig, Bremer in prep.) Thunberg’s description [Thunberg, 1822] of the Japanese tenebrionid beetle Hypophlaeus japonicus was found. This name cannot be found in the catalogues on Tenebrionidae [Gemminger, Harold, 1870; Gebien, 1910–1911, 1940; Löbl et al., 2008]. There are only two mentions of the species after the original description: Sherborn [1927] as Hypophloeus japonicus (incorrect subsequent spelling of the generic name) and Wallin, Wallin [2001] as Hypophlaeus japonicus.
The original description by Thunberg [1822: 7] is insufficient to identify the species:
“HYPOPHLOEUS japonicus: opacus, brunneus elytris striatis.
Minor Hyp. castaneo, major Hyp. depresso.
Magnitudine circiter Hypophl. bicoloris totus brunneus, opacus, planiusculus.
Elytra linearia, acuta, manifeste striata, abdomen obtegentia.
Similis valde et Hyp. depresso, sed major, obscurior, non nitidus, et duplo latior; et Hyp. fraxini, svecico, sed minime nitidus, verum opacus et elytra manifesta, etiam oculo nudo striata.”
The study of the syntype (Fig. 1–2), stored in Uppsala University, Museum of Evolution (UUZM, Sweden), shows the identity with Tribolium castaneum [Herbst, 1797: 282]: Tribolium castaneum (Herbst, 1797) = Hypophlaeus japonicus Thunberg, 1822, syn. n.
In accordance with the Recommendation 73F of the International Code of Zoological Nomenclature [1999] this
specimen (sex unknown) is designated as the lectotype. It is labelled as: “Uppsala Univ. Zool. Mus. Thunbergsaml. nr. 6516 Hypophlaeus japonicus Japon. TYP” (printed on
red cardboard, curator’s label), “Tribolium castaneum (Herbst) det. M. Lillig, 2015” (printed on white cardboard), “Hypophlaeus japonicus, LECTOTYPUS, des. M. Lillig, 2016” (printed on red cardboard) (Fig. 3).
Furthermore a specimen of “Hypophlaeus barthelemensis” is deposited in UUZM. The details on the label are: “UPSZEN 81947, Hypophlaeus barthelemensis Thunberg #6517, Skåp 6 Låda 06:03” (white cardboard, curator’s label). It is also Tribolium castaneum (Herbst, 1797) and appears to have never been described. The only published beetle name barthelemensis found in literature is Harpalus barthelemensis Sturm, 1826: 148 (“barthelemensis. Forström. Ins. Barthel.”) which is a nomen nudum.
My thanks are due to Dr Hans Mejlon (Uppsala University, Museum of Evolution, Zoology Section) for the loan, to Mr Edgar Müller (Saarwellingen, Germany) for providing the photo of the lectotype of Hypophlaeus japonicus and to two anonymous reviewers for their valuable comments.
Gebien H. 1910–1911. Pars. 15, 22 (1910), pars. 28, 37 (1911): Tenebrionidae, Trictenotomidae. In: Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Berlin: W. Junk. Vol. 18: 1–740.
Gebien, H. 1940. Katalog der Tenebrioniden. Teil 2. Mitteilungen der Münchner Entomologischen Gesellschaft. 30: 1061–1092.
Gemminger M., de Harold B. 1870. Catalogus Coleopterorum husque descriptorum synonymicus et systematicus Vol. 7. Tenebrionidae, Nilionidae, Pythidae, Melandryidae, Lagriidae, Pedilidae, Anthicidae, Pyrochroidae, Mordellidae, Rhipidophoridae, Cantharidae, Oedemeridae: München: E.H. Gummi: 1801–2179 + 11.
254 M. Lillig
Fig. 1–3. Hypophlaeus japonicus Thunberg, 1822, lectotype (length 4.1 mm) and labels. 1 – dorsal view; 2 – lateral view; 3 – labels.
Рис. 1–3. Hypophlaeus japonicus Thunberg, 1822, лектотип (длина 4.1 мм) и этикетки. 1 – вид сверху; 2 – вид сбоку; 3 – этикетки.
Herbst J. F. W. 1797. Natursystem aller bekannten in- und ausländischen Insekten, als eine Fortsetzung der von Büffonschen Naturgeschichte. Der Käfer VII. Theil. Berlin: Geh. Commerzien-Raths Pauli. XI + 346 p.
International Code of Zoological Nomenclature. Fourth Edition. International Trust of Zoological Nomenclature. 1999. London. xxix+306 p.
Löbl I., Merkl O., Ando K., Bouchard P., Egorov L. V., Iwan D., Lillig M., Masumoto K., Nabozhenko M., Novák V., Petterson R., Schawaller W., Soldati F. 2008. Family Tenebrionidae Latreille, 1802. In: Catalogue of Palaearctic Coleoptera (I. Löbl, A. Smetana eds.). Volume 5. Tenebrionoidea. Stenstrup: Apollo Books: 105–352.
Sherborn C. D. 1927. Index animalium sive index nominum quae ab A. D. MDCCLVIII generibus et speciebus animalium imposita sunt. Sectio secunda a kalendis ianuariis, MDCCCI usque ad finem decembris, MDCCCL. Part XIII. Index implicates - laminella. London: Longsmans, Green & Co. & British Museum: 3127–3392.
Sturm J. 1826. Catalog meiner Insecten-Sammlung. Erster Theil. Käfer.
Nürnberg: Verfasser. VIII + 277 pp., 4 pls.
Thunberg, C. P. 1822. Fauna japonica. (Dissertatio resp. O. Wernberg).
Upsaliae: Exudebant Regiae Academiae Typographi. 8 p.
Wallin S., Wallin H. 2001. Catalogue of type specimens. 1. C. P. Thunberg (1743-1828), Insecta (revised version 6). Uppsala University: Museum of Evolution, Zoology section. 66 p.