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Кавказский энтомол. бюллетень 12(2): 269–272 © CAUCASIAN ENTOMOLOGICAL BULL. 2016
1Russian Entomological Society, Krasnodar, Russia. E-mail: miroshnikov-ai@yandex.ru
2Sochi National Park, Moskovskaya str., 21, Sochi, Krasnodar Region 354002 Russia
1Русское энтомологическое общество, Краснодар, Россия
2Сочинский национальный парк, ул. Московская, 21, Сочи, Краснодарский край 354002 Россия
Key words: Coleoptera, Cerambycidae, Cerambycini, Dymasius, new or little-known species, Vietnam, India, Myanmar, Laos, Thailand, Indonesia.
Ключевые слова: Coleoptera, Cerambycidae, Cerambycini, Dymasius, новый и малоизвестные виды, Вьетнам, Индия, Мьянма, Лаос, Таиланд, Индонезия.
Abstract. A new species, Dymasius fedorenkoi sp. n., is described from Vietnam. A new combination is proposed: Dymasius ornatus (Gressitt et Rondon, 1970), comb. n. ex Derolus Gahan, 1891. New records are given of Dymasius lundbergi Hüdepohl, 1998 and D. mandibularis (Gahan, 1891) from Indonesia, inthesouthof Borneo, of D. ornatus in the northeast of India and of D. lineolatus Holzschuh, 2015 from Thailand, all considerably extending their distribution ranges. Some morphological features of Dymasius gilvago Holzschuh, 1999, D. lundbergi, D. ornatus and D. lineolatus are provided, the male and the female of the latter two species, respectively, being thereby described for the first time. The lectotype (male) and a paralectotype (female) of Dymasius plagiatus Gahan, 1906 are designated. Abundant colour pictures of all studied species are presented.
Резюме. Описан новый вид Dymasius fedorenkoi sp. n. из Вьетнама. Предложена новая комбинация: Dymasius ornatus (Gressitt et Rondon, 1970), comb. n. ex Derolus Gahan, 1891. Новые находки Dymasius lundbergi Hüdepohl, 1998 и D. mandibularis (Gahan, 1891) в Индонезии, на юге Борнео, D. ornatus на северо-востоке Индии и D. lineolatus Holzschuh, 2015 в Таиланде значительно расширяют их ареал. Приведены некоторые морфологические особенности Dymasius gilvago Holzschuh, 1999, D. lundbergi,
D. ornatus и D. lineolatus, причем у двух последних видов впервые описаны самец и самка соответственно. Обозначены лектотип (самец) и паралектотип (самка) Dymasius plagiatus Gahan, 1906. Представлено большое количество цветных иллюстраций всех рассматриваемых видов.
The genus Dymasius J. Thomson, 1864 is morphologically one of the most diverse and taxonomically very complex genera of the tribe Cerambycini Latreille, 1802. Its intrageneric systematics is still very poorly worked out, as the taxonomic attribution of a number of its formal representatives is highly controversial and cannot be definitively established. At the same time, some attempts at improving the taxonomy of Dymasius have recently been made. In particular, Holzschuh [2015] noted the genus as being highly heterogeneous, also confirming his own earlier considerations [Holzschuh, 1999] concerning the independence of the genus Microdymasius Pic, 1946 (described as a subgenus of Dymasius and treated of the same rank in various papers [Gressitt, Rondon, 1970; Holzschuh, 1984; Hüdepohl, 1998; Heffern, 2013; Nga et al., 2014; etc.]), proposed to refer only two species to this taxon, namely, M. angustatus (Pic, 1946) and M. niger (Gahan, 1906) (= Microdymasius honestus Holzschuh, 1999). However, presently among the unresolved problems such as dubious and highly disputable differences between Dymasius and Elydnus Pascoe, 1869 or between Dymasius and Derolus Gahan, 1891 are clearly notable. The results of some of my preliminary studies show that the distinctions between these taxa as specified in various publications [in particular, Gahan, 1906; Gressitt, 1951; Gressitt, Rondon, 1970; Hüdepohl, 1998; etc.] can only be used to some measure, but far from completely. In general, currently the formulation of clear-cut or reasonable differential diagnoses of these taxa (as well as of a number of others in the tribe Cerambycini) is very difficult.
270 A.I. Miroshnikov
The present paper describes a new species from Vietnam, provides new records of three little-known species from India and Indonesia, thereby considerably expanding their distribution ranges. The first description is given of the previously unknown female of a recently established species from Laos, and the lectotype and a paralectotype of a species from Myanmar are also designated. Even though the systematic position of one of the little-known species is not clear enough, on the basis of some of its features a new combination is proposed, the hitherto unknown male of that species being thereby described as well. Besides this, all species reviewed and illustrated below clearly demonstrate a remarkable morphological diversity of the genus Dymasius.
The material treated in this work belongs to the following institutional and private collections:
BM – Bishop Museum (Honolulu, USA);
BMNH – Natural History Museum (London, United Kingdom);
NHMD – Natural History Museum of Denmark, University of Copenhagen (Copenhagen, Denmark);
ZIN – Zoological Institute of the Russian Academy of Sciences (St. Petersburg, Russia);
ZSM – Zoologische Staatssammlung München (München, Germany);
cAM – collection of Alexandr Miroshnikov (Krasnodar, Russia);
cEV – collection of Eduard Vives (Barcelona, Spain); cLD – collection of Luboš Dembický (Brno, Czech
Republic);
cSM – collection of Sergey Murzin (Moscow, Russia).
Dymasius fedorenkoi Miroshnikov, sp. n.
(Color plate 7: 1–3)
Material. Holotype, ♂ (ZIN) (Color plate 7: 1): Vietnam, Kon Tum Prov., Kon Plong Distr., Dak Khe River, 14°43′20″N / 108°18′58″E, 1030 m, 8–23.04.2015, at light (leg. D. Fedorenko). Paratypes: 3♂ (cEV), Vietnam, Khanh Hoa Prov., Cam Lam Distr., Hon Ba Nature Reserve, 12°07ʹN / 108°57ʹE, 4.05.2013 (local collector).
Diagnosis. Based on the contrasting bichromous legs, this new species resembles Dymasius cos Holzschuh, 1998 (Color plate 7: 4), but by the structure of the male antennae, antennomere ratios, the sculpture of the pronotum it seems to be most similar to D. mandibularis (Gahan, 1891). However, D. fedorenkoi sp. n. differs clearly from both, as well as from all other congeners, by a peculiar coloration of the legs and, besides this, by the original combination of some other features. It is distinguished from D. cos in the longer antennae, the much longer last antennomere, the more deeply emarginate eyes, the elytra being slightly shorter and more strongly narrowed towards the apex, the structure of their apical sutural angle, and the clearly light recumbent setation of the dorsum. The new species differs from D. mandibularis by the continuous setation of the elytra devoid of any sign of longitudinal stripes, the structure of their apical sutural angle, and the seemingly more strongly developed pronotal recumbent setation. By the contrasting bicolour legs, D. fedorenkoi sp. n. can also be compared to D. flavimembris Hüdepohl, 1989, but differs very easily, except for the peculiar coloration of the legs, in many characters, including the structure and coloration of
the antennae, the peculiar sculpture of the pronotum, the structure of the elytra, etc.
width 7.1 mm. Coloration of integument, except for femora, combines black and black-brown tones; all femora, in contrast with completely black tibiae and tarsi, almost entirely orange, only at apex are darkened by narrow ring.
Head clearly longitudinal, with a well-expressed constriction behind temples, behind eyes dorsally with distinct punctures and small transverse wrinkles, and ventrally mostly with coarse transverse folds and partly with a heterogeneous coarse puncturation, predominantly in area of submentum; with well- developed antennal tubercles; eyes moderately convex, with a very deep emargination, their inner margins on ventral side very widely spaced, as in Color plate 7: 3; genae barely longer than 1st and 2nd segments of maxillary palpi combined; mandibles long, rounded angularly, sharpened apically, at inner margins without teeth or evident denticles; antennae very long, about 2.3 times as long as body; length ratio of antennomeres 1–11, 18 : 3 : 34 : 18 :
32 : 33 : 32 : 29 : 30 : 32 : 115; antennomere 1 without a cicatrix (apical carina), antennomeres 1–4 mostly with a coarse sculpture; apical external angle of antennomeres 5–10 subrectangular, slightly drawn laterad; last antennomere very long, about 3.6 times as long as penultimate one.
Pronotum clearly longitudinal, noticeably wider at base than at apex, from both base and apex relatively strongly broadened towards the middle, at apex with a sharp constriction; dorsally with coarse, mostly transverse, partly sinuous folds, more or less symmetrically placed on either side of midline.
Scutellum slightly transverse, strongly impressed dorsally, rounded at apex.
Elytra strongly narrowed towards apex, 2.6 times as long as humeral width; apical sutural angle extended into a long, flattened, almost vertically placed, ensiform tooth; with a small, dense puncturation.
Prosternum in apical part with coarse, transverse folds and punctures, in basal part with a more heterogeneous and less strongly expressed sculpture; prosternal process quite wide between procoxae, strongly broadened at apex, in lateral view rectangular at the very apex; mesosternal process between mesocoxae about
1.6 times as broad as prosternal process; metasternum and sternites (visible) with a small, dense puncturation; last (visible) sternite weakly emarginate apically.
Legs moderately long; femora not claviform, without a carina along each side; tarsomere 1 about equal to tarsomeres 2 and 3 combined.
Setation on dorsum mainly golden yellow, on venter predominantly greyish; elytra clothed with the most dense and continuous setation very strongly hiding both coloration and puncturation.
Variability. Paratypes males closely resembles the holotype. Body length 28–38 mm, humeral width 6–9 mm (according to Dr. Eduard Vives, Barcelona, Spain; personal communication).
Remarks. In the recent paper dedicated to new records of the tribe Cerambycini from Vietnam [Nga et al., 2014], figure 15 on page 431 seems to show this very new species, albeit misidentified as Dymasius maculatus Gressitt et Rondon, 1970.
Etymology. The new species honours my colleague and friend, Dr. Dmitry N. Fedorenko (Institute for Problems
of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia), who collected the holotype.
Dymasius lundbergi Hüdepohl, 1998 (Color plate 8: 5–7; Color plate 10: 17–22)
Dymasius (Microdymasius) lundbergi Hüdepohl, 1998: 213. Type locality: Ost-Malaysia, Sabah (according to the original
Figs 1–4. Dymasius J. Thomson, 1864.
1–3 – D. fedorenkoi sp. n., holotype, male; 4 – D. cos Holzschuh, 1998, holotype, male (after Holzschuh, 1998, but colour photograph, reproduced courtesy of Luboš Dembický). 1, 4 – habitus, dorsal view; 2 – head, dorsal view, and pronotum; 3 – head and anterior legs, ventral view, and prosternum.
Рис. 1–4. Dymasius J. Thomson, 1864.
1–3 – D. fedorenkoi sp. n., голотип, самец; 4 – D. cos Holzschuh, 1998, голотип, самец (по Holzschuh, 1998, но фотография цветная, предоставленная Л. Дембицким); 1, 4 – общий вид сверху; 2 – голова сверху и переднеспинка; 3 – голова, передние ноги снизу и простернум.
_11_Miroshnikov/Image_002.jpg)
Figs 5–10. Dymasius J. Thomson, 1864, habitus, dorsal view.
5–7 – D. lundbergi Hüdepohl, 1998: 5 – holotype, 6 – from Trus Madi Mts., Sabah, Malaysia, 7 – from Loksado env., S Kalimantan, Indonesia; 8–9 –
D. plagiatus Gahan, 1906: 8 – lectotype, 9 – paralectotype; 10 – D. mandibularis (Gahan, 1891) from Loksado env., S Kalimantan, Indonesia; 5, 7, 8 – males; 6, 9, 10 – females.
Рис. 5–10. Dymasius J. Thomson, 1864, общий вид сверху.
5–7 – D. lundbergi Hüdepohl, 1998: 5 – голотип, 6 – из гор Трас Мади, Сабах, Малайзия, 7 – из окрестностей Локсадо, Южный Калимантан, Индонезия; 8–9 – D. plagiatus Gahan, 1906: 8 – лектотип, 9 – паралектотип; 10 – D. mandibularis (Gahan, 1891) из окрестностей Локсадо, Южный Калимантан, Индонезия; 5, 7, 8 – самцы; 6, 9, 10 – самки.
_11_Miroshnikov/Image_003.jpg)
Figs 11–16. Dymasius J. Thomson, 1864, habitus, dorsal view.
11–12 – D. ornatus (Gressitt et Rondon, 1970), comb. n.; 13, 16 – D. gilvago Holzschuh, 1999 (13 – holotype after Holzschuh, 1999, but colour photograph, reproduced courtesy of Luboš Dembický); 14–15 – D. lineolatus Holzschuh, 2015; 11, 13, 14, 16 – males; 12, 15 – females.
Рис. 11–16. Dymasius J. Thomson, 1864, общий вид сверху.
11–12 – D. ornatus (Gressitt et Rondon, 1970), comb. n.; 13, 16 – D. gilvago Holzschuh, 1999 (13 – голотип, по Holzschuh, 1999, но фотография цветная, предоставленная Л. Дембицким); 14–15 – D. lineolatus Holzschuh, 2015; 11, 13, 14, 16 – самцы; 12, 15 – самки.
_11_Miroshnikov/Image_004.jpg)
Figs 17–25. Dymasius J. Thomson, 1864.
17–22 – D. lundbergi Hüdepohl, 1998; 23–24 – D. ornatus (Gressitt et Rondon, 1970), comb. n.; 25 – D. verticosus Holzschuh, 2010; 17–19, 23–25 –
pronotum; 20–22 – fragment of basal part of elytra before middle; 17, 19, 20, 22, 23 – males; 18, 21, 24, 25 – females.
Рис. 17–25. Dymasius J. Thomson, 1864.
17–22 – D. lundbergi Hüdepohl, 1998; 23–24 – D. ornatus (Gressitt et Rondon, 1970), comb. n.; 25 – D. verticosus Holzschuh, 2010; 17–19, 23–25 –
переднеспинка; 20–22 – фрагмент основной части надкрылий перед серединой; 17, 19, 20, 22, 23 – самцы; 18, 21, 24, 25 – самки.
A new species of the genus Dymasius J. Thomson, 1864 from Vietnam, with new data 271
description and the label of the holotype). Heffern, 2013: 10 (Borneo).
Material. Holotype, ♂ (ZSM) (Color plate 8: 5), Ost-Malaysia,
Sabah, 04.1994 (leg. Allen); 1♀ (NHMD), Malaysia, Sabah, Crocker Range, 03.2003 (local collector); 1♂ (NHMD), same, but 04.2006; 1♀ (cAM) (Color plate 8: 6), Malaysia, Sabah, Trus Madi Mt., 5°26′N / 116°27ʹE, 1000–1200 m, 17–29.04.2007 (leg. V. Tuzov); 1♂ (cAM ex collection of Alexey Klimenko, Tver, Russia), same, but 24.08.2012 (leg. A. Klimenko); 1♂ (cLD) (Color plate 8: 7), Indonesia, S Kalimantan, Kandangan Distr., 17 km NE of Loksado, 1000 m, 15.11.1997–15.01.1998 (leg. S. Jakl).
12.6 mm, humeral width 2.5 mm; the Indonesian male larger, 16.2 mm and 3.3 mm, respectively. In Dymasius lundbergi, pattern of pronotum’s dorsum (formed by sculpture and light setation) somewhat variable, as in Color plate 10: 17–19; puncturation of a peculiarly looking longitudinal fragment on elytral disk before middle on each side of suture (devoid of setation) in the Indonesian male mainly relatively slightly sparser and larger than in Malaysian males (including the holotype) (Color plate 10: 20–22), this making puncturation look more strongly expressed.
Distribution. This species was described from three specimens from Sabah, Malaysia, thereby the holotype (male) and one paratype (female) were accompanied by no detailed locality data, whereas the provenance of the remaining paratype (male) was clarified: “Kumanis Road, 10th mile” [Hüdepohl, 1998].
Based on the studied material, D. lundbergi is being recorded here in Indonesia (including the south of Borneo) for the first time.
Dymasius ornatus (Gressitt et Rondon, 1970), comb. n.
(Color plate 9: 11, 12; Color plate 10: 23, 24)
Derolus ornatus Gressitt et Rondon, 1970: 76. Type locality: km 17 of Paksane Road, Vientiane Prov., Laos (according to the original description). Nga et al., 2014: 432 (NE Vietnam).
Material. Holotype, ♀ (BM) (photograph by Nobuo Ohbayashi),
“(Bishop 8297), Km 17 Paksane Rd, 170 m, Vientiane Prov., Laos, 18.08.1963” / “Rondon coll.”; 1♂ (cLD) (Color plate 9: 11), 1♀ (cLD) (Color plate 9: 12), NE India, Meghalaya, 3 km E of Tura, 25°30′N / 90°14′E, 500– 1150 m, 15–22.04.1999 (leg. Dembický, Pacholátko).
Remarks. This species is one of the typical examples when currently no exact systematic position of a taxon can be determined. At the same time, although it has been described in the genus Derolus, I am inclined to transfer it to Dymasius due to certain features, especially those in the structure of the pronotum. The pronotum of the holotype (female) and that in the Indian female are slightly longitudinal (noted in the original description of this species), in the Indian male even more longitudinal, thereby the setation of the pronotum’s dorsum in all three specimens is well-developed (including that on the disk) and forms a clear peculiar pattern (Color plate 10: 23, 24) which is typical of a number of Dymasius, in contrast to most of Derolus. By the shape, sculpture and character of setation of the pronotum, D. ornatus seems to be quite similar to D. verticosus Holzschuh, 2010 (Color plate 10: 25), D. nimbatus Holzschuh, 1991 and some other species. Although tarsomere 1 in D. ornatus is shorter than tarsomeres 2 and 3 combined, while the femora and tibiae each are with a clear carina along each side, as is typical of Derolus, these features are also observed in different Dymasius species.
Based on all above evidence, the new combination is established: Dymasius ornatus (Gressitt et Rondon, 1970), comb. n. (from Derolus).
This species was described from two females [Gressitt, Rondon, 1970], the male hitherto remaining unknown.
Body length of the studied female (Color plate 9: 12),
13.8 mm, humeral width 3 mm. The elytra in both Indian specimens are more strongly elongated than in the holotype.
Distribution. This species was described from northern Laos. It has recently been recorded from the northeast of Vietnam [Nga et al., 2014].
Based on the studied material, D. ornatus is being recorded here from India for the first time.
Dymasius plagiatus Gahan, 1906 (Color plate 8: 8, 9)
Dymasius plagiatus Gahan, 1906: 141. Type locality: Burma (now Myanmar), Karen Mts (according to the original description and the label of the lectotype). Aurivillius, 1912: 60.
Material. Lectotype, ♂, here designated (BMNH) (Color plate 8: 8),
“Birmah, Karen Mts” / “Doherty” / “Fry Coll. 1905–100.” / “Dymasius plagiatus Gahan Type” / “Type” + “Lectotypus ♂ Dymasius plagiatus Gahan, 1906, A. Miroshnikov des., 2016”; paralectotype, ♀, here designated (BMNH) (Colo plate 8: 9), “Birmah, Karen Mts” / “Doherty” / “Fry Coll. 1905–100.” + “Paralectotypus ♀ Dymasius plagiatus Gahan, 1906,
A. Miroshnikov des., 2016”.
Remarks. This species was described from one male and one female [Gahan, 1906], hitherto remaining known only from those two type specimens. In the original description, the body size was not indicated separately for the male and the female. In the present paper, Dymasius plagiatus is illustrated for the first time.
Dymasius lineolatus Holzschuh, 2015 (Color plate 9: 14, 15)
Dymasius lineolatus Holzschuh, 2015: 45. Type locality: Laos, Luang Phrabang env., 19°53.420′N / 102°08.229′E (according to the original description).
Material. 3♂, 1♀ (cLD), 1♂ (cAM ex cLD), Laos, Luang Phrabang
env., 19°53.420′N / 102°08.229′E, 16–19.02.2010 (leg. M. Pejcha); 2♂, 4♀
(cSM), 2♀ (cAM ex cSM), N Thailand, 100 km N Chaing Mai, Chiang Dao Hill Resort, 600 m, 10–23.03.2010 (leg. S. Murzin).
Morphological notes. Female (Color plate 9: 15). Closely resembling the male. Body length 10.7–12.6 mm, humeral width 2.15–2.7 mm. In comparison with the male, antennae shorter, slightly shorter than body, elytra more strongly narrowed towards apex.
272 A.I. Miroshnikov
In some males, longitudinal stripes of dense, recumbent, golden setae forming a peculiar pattern on pronotum, the stripes being more strongly developed than in holotype, clearly wider; thereby two symmetrical paramedian stripes being complete, extending from apex to pronotum base (Color plate 9: 14). Elytral pattern slightly variable. Body length of studied specimens, 9.9–13.3 mm, humeral width 2.1–2.9 mm.
D. lineolatus is being recorded here from Thailand for the first time.
Dymasius gilvago Holzschuh, 1999 (Color plate 9: 13, 16)
Dymasius gilvago Holzschuh, 1999: 22. Type locality: S India, Tamil Nadu, Nilgiri hills, 15 km SE of Kotagiri, near Kunchappanai, 11°22′N / 76°56′E, 900 m (according to the original description).
Material. 1♂ (BMNH) (Color plate 9: 16), [? S India] “46/6” /
“Sebasmia?” + “Dymasius gilvago Holzschuh, 1999 ♂ A. Miroshnikov det., 2016”; holotype, ♂ (collection of C. Holzschuh, Villach, Austria) (Color plate 9: 13; photograph by L. Dembický), S India, Tamil Nadu, Nilgiri hills, 15 km SE of Kotagiri, near Kunchappanai, 11°22′N / 76°56′E, 900 m, 13– 20.05.1994 (leg. Z. Kejval, R. Sauer).
Remarks. Thisspecieswasdescribedfromasinglemale [Holzschuh, 1999]. The male referred to here is the second specimen of D. gilvago known to me (Color plate 9: 16), but it lacks a geographical label. Perhaps, like the holotype, it also comes from southern India. Its body size is barely smaller than the holotype: length 14.4 mm, humeral width
3.2 mm (according to the original description, the body of the holotype is 15.1 mm long). This male closely resembles the holotype (Color plate 9: 13) and its pronotum shows nearly the same peculiar pattern of dense, recumbent, light setae and a similar sculpture of transverse, sharp folds, while the elytra support the same monochromatic, almost continuous, dense setation.
Dymasius mandibularis (Gahan, 1891) (Color plate 8: 10)
Imbrius ? mandibularis Gahan, 1891: 21. Type locality: [W Malaysia] Penang (according to the original description).
Imbrius mandibularis: Aurivillius, 1912: 60.
Dymasius mandibularis: Holzschuh, 2005: 9.
Dymasius (Dymasius) mandibularis: Heffern, 2013: 9.
Dymasius obscurus Hüdepohl, 1998: 212. Type locality: Ost Malaysia, Sabah, Crocker Range (according to the original description and the label of the holotype). Holzschuh, 2005: 9 (syn. pro D. mandibularis).
Material. 1♀ (cLD) (Color plate 8: 10), Indonesia, S Kalimantan,
Kandangan Distr., 17 km NE of Loksado, 1000 m, 15.11.1997–15.01.1998 (leg. S. Jakl).
Distribution. This species has hitherto been known only from Malaysia, both the western and eastern parts [Gahan, 1891; Hüdepohl, 1998].
Based on the studied material, D. mandibularis is being recorded here in Indonesia (including the south of Borneo) for the first time.
I am most grateful to Dmitry N. Fedorenko, who was funded by the Russia-Vietnam Tropical Center, for the
valuable material he rendered to me for study. My sincere thanks also go to Michael Balke and Katja Neven (ZSM), Maxwell V.L. Barclay and Michael F. Geiser (BMNH), Alexey Yu. Solodovnikov and Sree Gayathree Selvantharan (NHMD) for the opportunity to study the museum material under their care while Luboš Dembický (Brno, Czech Republic), Sergey V. Murzin (Moscow, Russia) and Eduard Vives (Barcelona, Spain) have provided some specimens (or the pictures of the specimens) from their private collections. I am deeply indebted to Kirill V. Makarov (Moscow Pedagogical State University, Russia) who has helped with the preparation of photographs, again to Luboš Dembický for the pictures of the holotypes of many Dymasius species, including D. cos, to Nobuo Ohbayashi (Kamimiyada, Miura City, Japan) for the photograph of the holotype of Dymasius ornatus, to Alexey A. Klimenko (Tver, Russia), again Luboš Dembický and Sergey V. Murzin who have generously presented me with some specimens of Dymasius spp., to Francesco Vitali (Luxembourg) and Sergi Trócoli (Barcelona, Spain) for the valuable information.
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