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Кавказский энтомол. бюллетень 11(2): 385–388 © CAUCASIAN ENTOMOLOGICAL BULL. 2015
A.N. Severtzov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect, 33, Moscow 119071 Russia. E-mail: danilevskyml@rambler.ru, danilevsky@cerambycidae.net
Институт проблем экологии и эволюции им. А.Н. Северцова РАН, Ленинский пр., 33, Москва 119071 Россия
Key words: Coleoptera, Cerambycidae, Cerambycinae, Purpuricenus, new species, Caucasus, Armenia, Azerbaijan, Georgia, Russia.
Ключевыеслова: Coleoptera, Cerambycidae, Cerambycinae, Purpuricenus, новый вид, Кавказ, Армения, Азербайджан, Грузия, Россия.
Abstract. Purpuricenus caucasicola sp. n. is described from Krasnodar Region of Russia. The species is also distributed in Transcaucasia (Azerbaijan, Georgia) and probably in North Turkey. The taxon was recorded for the region before with a wrong name “Purpuricenuscaucasicus”. All similar taxa are also accepted as species: P. graecus Sláma, 1993, P. renyvonae Sláma, 2001, P. baeckmanni Danilevsky, 2007, P. neocaucasicus Rapuzzi et Sama, 2013. Резюме. Purpuricenus caucasicola sp. n. описан из Краснодарского края России. Вид также распространен в Закавказье (Азербайджан, Грузия) и, вероятно, в Северной Турции. Таксон ранее указывался для региона с неверным названием «Purpuricenus caucasicus». Все близкие таксоны также рассматриваются как виды: P. graecus Sláma, 1993, P. renyvonae Sláma, 2001,
P. baeckmanni Danilevsky, 2007, P. neocaucasicus Rapuzzi et Sama, 2013.
The name Purpuricenus budensis var. caucasicus Th. Pic, 1902 [Th. Pic, 1902] was accepted [Heyden et al., 1906: 517; М. Pic, 1912: 13) as an aberration of P. budensis (Götz, 1783). That status was proved by Plavilstshikov [1940: 568–569], who also described P. budensis m. productus Plavilstshikov [1940: 569] – unavailable name. Both forms were figured by Plavilstshikov [1940: 568–569]. Miroshnikov [1984: 7, 9] identified a species from the Caucasus and Crimea as P. caucasicus and included in the taxon P. budensis m. productus. The reasons for such identification were not published by him, but evidently it was made on the base of Plavilstshikov’s [1940: 568–569] pictures and specimens from Plavilstshikov’s collection without investigation of Th. Pic’s type.
Purpuricenus caucasicus sensu Miroshnikov [1984] was accepted by Danilevsky and Miroshnikov [1985], Sláma [1993, 2001], Sláma and Slámová [1996], Althoff and Danilevsky [1997], Sama [2002], Özdikmen and Çaglar [2004].
Later the species was divided in two subspecies with the separation of Crimean populations as P. c. baeckmanni
Danilevsky, 2007 [Danilevsky, 2007: 38]. That taxon was originally segragated by J. Baeckman [see “Purpuricenus budensus tauricus Baeckm. in litt.” by Plavilstshikov, 1940: 569], but not described. Sama [2010: 52] accepted the segregation of Crimean subspecies from Caucasian subspecies, but paradoxally joined it with Balcanian subspecies publishing new synonyms: P. caucasicus renyvonae Slama, 2001 = P. c. baeckmanni Danilevsky, 2007. The natural situation was restored by Danilevsky [2010: 231] with separate publication of P. c. baeckmanni for Ukraine and P. caucasicus renyvonae for Bulgaria, Greece, Macedonia and Yugoslavia. It was supported by Miroshnikov [2009b: 64; 2012: 45] with atribution of the nominative subspecies to the Caucasus and Turkey.
The holotype of P. b. var. caucasicus was investigated by Sabbadini and Pesarini [1992], who agreed with the original position of the name inside P. budensis and published “Purpuricenus budensis caucasicus” as a Transcaucasian subspecies also distributed in allied territories of Turkey. The authors published the label of the holotype (“Helendorf”), but wrongly applied that old name to Armenian city Sevan. The elytral design of the holotype of P. b. var. caucasicus was published much earlier by Villiers [1978: 315, Fig. 1073], who also kept it inside
P. budensis. The real nature of the holotype as P. kaehleri (Linnaeus, 1758) was identified by Rapuzzi and Sama [2013: 158], but the location of that specimen (“Helendorf”) was wrongly determined (following Sabbadini and Pesarini) as “Elenovka, now Sevan”. The true Helendorf was shown before by Miroshnikov [2012a: 45]: Khanlar (now Göygöl) in Azerbaijan.
The Caucasian species P. caucasicus sensu Miroshnkov [1984] (not Th. Pic, 1902) was joined by Rapuzzi and Sama [2013: 156, part.] to their P. renyvonae ssp. neocaucasicus Rapuzzi et Sama, 2013 distributed from Bolu, Kastamonu, Çorum and Tokat (Turkey) to the Caucasus (on the base of a single male from Tbilisi). So, Caucasian populations rested unknown to Rapuzzi and Sama, while Turkish populations were unknown to M. Danilevsky in all his publications.
386 M.L. Danilevsky
In fact many Purpuricenus taxa of the group are poorly investigated and known in a few specimens only. All are allopatric and usually more or less isolated, that allows to fabricate intentionally species taxa by selective joining similar populations. The modern level of knowledge permits to upgrade most of available names to species rank: P. graecus Sláma, 1993, P. renyvonae Sláma, 2001,
P. baeckmanni Danilevsky, 2007, P. neocaucasicus Rapuzzi et Sama, 2013.
The Caucasian species is described bellow as new. Abbreviations of collection:
AM – collection of A. Miroshnikov (Krasnodar, Russia);
JV – collection of J. Vartanis (Uherský Brod, Czech Republic);
NMP – National Museum Prague (Czech Republic);
ZMM – Zoological Museum of Moscow University (Russia).
Purpuricenus caucasicola sp. n.
(Color plate 11: 1–4)
Purpuricenus (s. str.) budensis ab. caucasicus: Plavilstshikov, 1931: 80 – Caucasus.
Purpuricenus budensis ab. caucasicus: Plavilstshikov, 1940: 568–569, 769, part.; Heyrovský, 1955: 241, obr. 44 – 4; Panin,
Săvulescu, 1961: 365; Kaszab, 1971: 202–203.
Purpuricenus budensis morpha productus: Plavilstshikov, 1940: 568–569, 769, part. (not available name) – Crimea, Caucasus, Transcaucasia, Syria; 1948: 118 – Transcaucasia; Heyrovský, 1955:
241, obr. 44 – 9–10; Panin, Săvulescu, 1961: 365.
Purpuricenus budensis ab. productissimus: Plavilstshikov, 1940: 569, 769 (not available name); Heyrovský, 1955: 241, obr.
44 – 10; Panin, Săvulescu, 1961: 365.
Purpuricenus budensis rasa productus: Panin, Săvulescu, 1961: 365 – “Crimeea, Caucaz, Transcaucazia şi Siria”.
Purpuricenus budensis f. productus: Demelt, 1968: 65, 67, part. – “Krim, Kaukasicus, Transkaukasien”; Bernhauer, 1976: 124–125.
Purpuricenus budensis f. budensis ab. caucasicus: Demelt, 1968: 66, part.
Purpuricenus budensis f. productus ab. productissimus: Demelt, 1968: 67, part.
Purpuricenus budensis var. productus: Villiers, 1978: 315,
part.
Purpuricenus budensis var. productissimus: Villiers, 1978: 315, part.
Purpuricenus caucasicus: Miroshnikov, 1984: 7, 9 – North- West Caucasus; 2000: 233 – Crimea, ? South and Central Europe, Adygea; 2007: 182, part. – Turkey (to the east from Kastamonu), Crimea, Caucasus (Kislovodsk, Gori, Mtskheta, Lagodekhi), Krinitsa environs in Krasnodar Region; 2009a: 791 – Belaya River valley in Adygea; 2009b: 64, part. – Crimea, Asia Minor, Caucasus (nominative subspecies): Khamyshki; 2012a: 44, part. – South Crimea, Asia Minor, Caucasus, ? Syria, ? North-West Iran; 20132b: 291, part. – South Crimea, Asia Minor, Caucasus; Danilevsky, Miroshnikov, 1985: 212–214, part. – Crimea, Caucasus, ? South and Central Europe; Sláma, 1993: 56; 2001: 225-227, 235-237, part.; Sláma, Slámová, 1996: 123, part. – Yug. – Makedonia; Althoff, Danilevsky, 1997: 21, part. – ? South Europe (including Crimea); Zamotailov, Miroshnikov, 1997: 181; Sama, 2002: 55; 2010: 52,
part.; Özdikmen, 2007: 247, 389, part. – “Europe (Crimea), Caucasus, Turkey [including Abant, Erzurum, Muğla and Bolu]”; 2008: 371; Nikitsky et al., 2008: 343 – Krasnodar Region; Bartenev, 2009: 175 – South and Central Europe, Greece, Bulgaria, Romania, Ukraine (Crimea), ? Caucasus; Özdikmen, Çaglar, 2004: 57 –
Turkey; Nieto, Alexander, 2010: 31 – Europe; Sakenin et al., 2011: 5 – “Iran: East Azarbayjan province: Shabestar.”
Purpuricenus budensis caucasicus: Sabbadini, Pesarini, 1992: 56, part. (the holotype label: “Helendorf”) – Armenia, Turkey.
Purpuricenus caucasicus caucasicus: Danilevsky, 2007: 36 – “Caucasus with Transcaucasia, Anatolia” (from Erzurum to Bolu); 2010: 231 – Azerbaijan, Armenia, Georgia, Turkey, South Russia; Miroshnikov, 2012a: 45 – nominative subspecies is distributed in the Caucasus and Asia Minor.
Purpuricenus budensis ? caucasicus: Özdikmen et al., 2009: 75.
Purpuricenus budensis productus: Özdikmen et al., 2009: 75 – “S Turkey”.
Purpuricenus (s. str.) caucasicus caucasicus: Catalogue..., 2010: 198 – Azerbaijan, Armenia, Georgia, Turkey, South Russia; Miroshnikov, 2010: 249 – Adygea (Khamyshki).
Purpuricenus (s. str.) caucasicus: Miroshnikov, 2011: 558 –
? Novorossiysk; 2012: 42 (Caucasian areal map), 44–45, Pl. 5.
Purpuricenus renyvonae ssp. neocaucasicus Rapuzzi et Sama, 2013: 156, part. – from Bolu, Kastamonu, Çorum and Tokat (Turkey) to the Caucasus (1 male) (Georgia).
Material. Holotype, ♂: NW Caucasus, Novorossiysk env., Dyurso,
Orel Mt., 30.06–13.07.2009, A. Miroshnikov leg. (ZMM). Paratypes: 1♂ with 4 labels: “Geok-Tapa, Caucasus, A. Schelkownikow”, “Cotypus”, “Purpuricenus budensis Gz. m. productus m. N. Plavilstshikov det.”, “Purpuricenus caucasicus Pic ♂ A. Miroshnikov det. 1984” (ZMM); 1♀, “Caucas. bor. Kislovodsk” (ZMM); 1♀ with 4 labels: “Transcauc. Lagodechi, 9.07.[1]913, Mlokoss.”, “Cotypus”, “Purpuricenus budensis Gz. m. productus
N. Plavilstshikov det.”, “Purpuricenus caucasicus Pic ♀ A. Miroshnikov det. 1984” (ZMM); 1♀, “Лагодехи, 17.07.1926, Млокосевич” (Lagodekhi, 17.07.1926, Mlokosevich) (AM); 1♀, “prov. Kuban, fl. Belaja, Chamyshki, 6.08.[19]33, Arnoldi” (ZMM); 1♀, Dzhubga, Krinitsa, 8.07.1975,
Zamotailov leg. (AM); 1♂, Georgia, Tsagveri, 15.07.1981, V. Dolin leg. (AM); 1♂, “Gruzia, Tbilisi, Jezero Lisy [Lisi Lake], 29.6.1983, Dalihod leg.” (JV); 1♂, 1♀: NW Caucasus, Novorossiysk env., Dyurso, Orel Mt., 30.06– 13.07.2009, A. Miroshnikov leg. (AM); 1♀, NW Caucasus, Novorossiysk env., Malyi Utrish, 30.06–13.07.2009, A. Miroshnikov leg. (AM); 5♂, 4♀, the same locality, 29.06–18.07.2011, A. Miroshnikov leg. (AM); 2♂, 2♀, NW Caucasus, Anapa env., Bolshoy Utrish, 20.06.2012, G. Shemberger leg. (AM); 1♂, Krasnodar Region, 17 km south of Anapa, 2.07.2012,
G. Shemberger leg. (JV); 1♀, Krasnodar Region, 17 km S.O. Anapa, 30.06.2014 (JV).
Purpuricenus neocaucasicus Rapuzzi et Sama, 2014: 1♂, 1♀ with labels: “Abant (Bolu), TR, 7.7.1994, col. Auvray N.” (AM); 1♀ (Color plate : 5) with a label: “Turkva-Yaraligöz, Kastamonu prov., 10.7.2008, D.Loupanec leg.” (JV);1♂ with a label: “Yaraligoz TR (Kastamonu), 15.07.1996, leg. Auvray N.” (NMP); 1♂ with a label: “Abant (Bolu) TR, 20.07.2000, leg. Auvray N.” (NMP).
Description. Body relatively narrow; antennae in males usually surpassing elytra by 3rd apical joints, often by 2nd, but sometimes by 1st only; antennae in females usually reaching last elytral forth, sometimes hardly reaching last elytral third, very rare nearly attain elytral apex; 4th antennal joint as long as 5th or a little shorter, in males much longer than 1st or in females about equal to 1st, and much shorter than 3rd in males and females; thorax transverse, in males about as long as basal width or a little shorter; in females usually shorter than basal width, very rare about equal; lateral thoracic tubercles in males short, more or less obliterated, in females short, but usually distinct; prothorax convex, always with small shining central tubercle; basal pronotal half in males usually black up to central tubercle, very rare wider, only two males (Orel Mt. in Dyurso env. and Lisi Lake near Tbilisi) with about totally black pronotum bearing small reddish spots; pronotum in females much more red; posterior black area rarely reaching central tubercle; scutellum elongated; elytra parallelsided, in males from 2.3 to 2.5 times longer than basal width; in females from 2.1 to 2.3 times; black elytral area rather long, more or less parallelsided, slightly widened at apex, sometimes (1 male from Lisi Lake and 1 female from Lagodekhi) touching scutellum, anteriorly protruding and rounded, very rare emarginated; elytral apices can be widely rounded (especially in females), but often more or less truncated with distinct external angles; pygidium in males and in
A new species of the genus Purpuricenus Dejean, 1821 (Coleoptera: Cerambycidae) from the Caucasus Color plate 11.
Figs 1–5. Purpuricenus, general view.
1–4 – P. caucasicola sp. n.: 1 –male, holotype, Novorossiysk env., Dyurso, Orel Mt. (photo by A.I. Miroshnikov and S.O. Kakunin); 2 – female, paratype from the same locality; 3 – male, paratype, Azerbaijan, Geok-Tapa; 4 – female, paratype, Georgia, Lagodekhi; 5 – P. neocaucasicus, female, Turkva-Yaraligöz, Kastamonu Prov., 10.7.2008, D. Loupanec leg.
Рис. 1–5. Purpuricenus, общий вид.
1–4 – P. caucasicola sp. n.самец, голотип, окрестности Новороссийска, Дюрсо, гора Орел (фото А.И. Мирошникова и С.О. Какунина); 2 – самка, паратип из того же места; 3 – самец, паратип, Азербайджан, Геок-Тапа; 4 – самка, паратип, Грузия, Лагодехи; 5 – P. neocaucasicus, самка, Turkva- Yaraligöz, Kastamonu Prov., 10.7.2008, D. Loupanec leg.
A new species of the genus Purpuricenus Dejean, 1821 (Coleoptera: Cerambycidae) from the Caucasus 387
females slightly emarginated, postpygidium truncated; body length in males 12–15.8 mm, in females 10.4–17.5 mm; body width in males 3.7–4.6 mm, in females 3.2–5.8 mm.
P. neocaucasicus (Color plate 11: 5) distributed in Western and Central Anatolia, which is known to me after original description and 5 specimens only. I was not able to get more specimens. So, the distinguishing characters need to be better studied: in general P. caucasicola sp. n. is distinctly narrower, with more red pronotum; anterior margin of black elytral area is rounded, very rare emarginated.
Anapa (Dyurso env., Orel Mt., 44°41′55″N / 37°31′50″E; Malyi Utrish Cape, 44°42′29″N / 37°27′30″E) to Kislovodsk and most probably further eastwards; another known localities are: Krinitsa (44°23′N / 38°20′43″E), Khamyshki (600 m, 44°6′N / 40°8′E). Georgia: from Borzhomi and Tsagveri to Lagodekhi; another known localities are: Gori, Mtskheta, Khashmi (41°46′N / 45°11′E). Azerbaijan: from Belokany (41°44′N / 46°25′E) to Gyandzha and Agdash (40°39′N / 47°28′E).
because such publications as Panin and Săvulescu [1961] with “budensis rasa productus Plav.” or Demelt [1968] of “budensis f. productus Plav.” both with exact own separate areas are not more that publications of real subspecies. But ICZN [1999] from one side does not allow regarding old unavailable names newly published as “forma” available, and from other side does not mention “rasa” names at all (neither “natio”). The early publication of the name “productus Plav.” (before 1985 – Art. 45.6.4.1.) as valid could make it available. But following explicitly to the Code it is necessary now to regard it unavailable.
I am very grateful to my friends and colleagues for providingmewiththespecimensforstudy: Aleksey Gusakov (Zoological Museum of Moscow University, Russia), Denis Kasatkin (Rostov-on-Don, Russia), Alexander Miroshnikov (Krasnodar, Russia), Janis Vartanis (Uherský Brod, Czech Republic).
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